Table 4.
Morphological and meristic data of C. alpinusFatio 1885 from lakes Thun and Brienz, MHNG-717.045 lectotype from Lake Thun; non-type material N = 21 from Lake Thun and N = 9 from Lake Brienz.
| Morphological characters | C. alpinus | Both lakes | Lake Thun | Lake Brienz | |||||||||||
| Lectotype | Non-types both sexes | Non-types | Non-types | ||||||||||||
| N-total = 30 | N-total = 21 | N-females = 12 | N-males = 9 | N-total = 9 | N-females = 6 | N-males = 3 | |||||||||
| Mean ± StDev | Range | Mean ± StDev | Range | Mean ± StDev | Range | Mean ± StDev | Range | Mean ± StDev | Range | Mean ± StDev | Range | Mean ± StDev | Range | ||
| SL (mm) | 283.0 | 266.1±56.4 | (147-364) | 288.3±45.1 | (210-364) | 299.6±37.3 | (267-364) | 273.3±52.2 | (210-352) | 214.3±46.0 | (147-290) | 240.1±27.9 | (213-290) | 162.7±23.3 | (147-190) |
| Percentage of standard length | |||||||||||||||
| PelvFB | 4.5 | 4.2±0.4 | (3.1-5.0) | 4.4±0.3 | (3.8-5) | 4.3±0.4 | (3.8-5.0) | 4.4±0.3 | (4.0-4.8) | 3.7±0.3 | (3.1-4.0) | 3.9±0.1 | (3.7-4.0) | 3.5±0.5 | (3.1-4.0) |
| PelvFS | 4.7 | 6.1±0.8 | (4.7-7.6) | 5.9±0.7 | (4.7-7.2) | 5.9±0.7 | (4.8-6.6) | 5.9±0.8 | (4.7-7.2) | 6.7±0.5 | (5.9-7.6) | 6.8±0.5 | (6.0-7.6) | 6.5±0.6 | (5.9-7.1) |
| PelvF | 18.1 | 16.7±1.1 | (14-18.2) | 16.6±0.9 | (14.7-18.2) | 16.2±0.9 | (14.7-17.3) | 17.2±0.6 | (16.3-18.2) | 16.9±1.5 | (14-18.1) | 16.9±1.6 | (14-18.1) | 16.8±1.4 | (15.2-18) |
| PecFB | 3.7 | 3.3±0.3 | (2.5-3.7) | 3.4±0.3 | (2.8-3.7) | 3.4±0.3 | (2.8-3.7) | 3.3±0.2 | (3.0-3.5) | 3.0±0.3 | (2.5-3.5) | 3.0±0.3 | (2.5-3.5) | 3.1±0.3 | (2.8-3.3) |
| PecF1 | 18.7 | 16.2±1.2 | (13.6-18.7) | 16.2±1.3 | (13.6-18.7) | 16.0±1.4 | (13.6-18.6) | 16.5±1.1 | (15.1-18.7) | 16.3±1.1 | (13.9-17.1) | 16.4±1.3 | (13.9-17.1) | 16.1±0.9 | (15.1-16.8) |
| PecF2 | 19.7 | 17±1.1 | (14.4-19.7) | 17±1.1 | (15.3-19.7) | 16.8±1.1 | (15.3-19.1) | 17.3±1.1 | (15.8-19.7) | 16.9±1.2 | (14.4-17.7) | 17±1.3 | (14.4-17.7) | 16.7±1.1 | (15.5-17.6) |
| DFB | 12.9 | 12.8±0.9 | (10.7-14.7) | 12.9±0.9 | (11.5-14.7) | 12.9±0.9 | (11.7-14.4) | 12.9±0.9 | (11.5-14.7) | 12.6±1.1 | (10.7-14.3) | 12.6±1.1 | (10.7-14.0) | 12.7±1.4 | (11.7-14.3) |
| DFAe | na | 19.7±1.2 | (16.4-23.0) | 19.5±0.8 | (17.6-20.6) | 19.3±0.8 | (17.6-20.5) | 19.7±0.8 | (18.6-20.6) | 20.3±1.8 | (16.4-23.0) | 20.1±1.9 | (16.4-21.5) | 20.9±1.9 | (19.5-23.0) |
| DFAd | 22.6 | 20.8±1.4 | (17.0-24.0) | 20.6±0.9 | (18.6-22.5) | 20.4±0.8 | (18.6-21.3) | 20.9±1.1 | (19.6-22.5) | 21.4±2.0 | (17.0-24.0) | 21.2±2.2 | (17.0-22.7) | 21.7±2.0 | (20.0-24.0) |
| DFPe | 5.6 | 5.0±0.6 | (4.0-6.5) | 4.8±0.5 | (4-5.8) | 4.8±0.6 | (4.0-5.8) | 4.9±0.4 | (4.2-5.5) | 5.3±0.7 | (4.3-6.5) | 5.2±0.6 | (4.3-5.9) | 5.5±1.1 | (4.4-6.5) |
| AFB | 12.5 | 12.4±0.9 | (9.8-14.2) | 12.5±0.9 | (10.3-14.2) | 12.3±0.9 | (10.3-14.0) | 12.8±0.8 | (11.1-14.2) | 12.1±1.0 | (9.8-13.3) | 12.1±1.1 | (9.8-12.9) | 12.2±1.0 | (11.4-13.3) |
| AFAe | 14 | 12.3±0.9 | (9.8-13.8) | 12.2±0.7 | (10.5-13.3) | 11.9±0.7 | (10.5-12.7) | 12.5±0.7 | (10.9-13.3) | 12.5±1.3 | (9.8-13.8) | 12.6±1.4 | (9.8-13.8) | 12.4±1.3 | (11.2-13.7) |
| AdFB | 4.8 | 4.4±0.6 | (3.4-5.5) | 4.3±0.6 | (3.4-5.5) | 4.3±0.6 | (3.5-5.5) | 4.2±0.5 | (3.4-4.7) | 4.7±0.5 | (3.8-5.4) | 4.6±0.5 | (3.8-5.1) | 4.9±0.5 | (4.5-5.4) |
| CF | na | 24±1.4 | (19.1-26.3) | 24±1 | (21.9-26) | 23.8±0.7 | (22.7-24.8) | 24.3±1.2 | (21.9-26) | 24.1±2.2 | (19.1-26.3) | 24±2.6 | (19.1-26.3) | 24.3±1.5 | (22.9-25.9) |
| CD | 8.8 | 8.1±0.5 | (6.7-8.9) | 8.2±0.4 | (7.6-8.9) | 8.1±0.3 | (7.6-8.6) | 8.3±0.4 | (7.8-8.9) | 7.8±0.6 | (6.7-8.6) | 7.8±0.6 | (6.7-8.6) | 7.8±0.8 | (7-8.4) |
| CL | 12.6 | 12.6±0.8 | (11.2-14.4) | 12.6±0.9 | (11.2-14.4) | 12.5±0.8 | (11.2-13.6) | 12.9±0.9 | (11.8-14.4) | 12.5±0.9 | (11.3-13.9) | 12.5±0.8 | (11.3-13.6) | 12.5±1.2 | (11.5-13.9) |
| PAdC | 18.4 | 17.7±1.1 | (15.3-19.5) | 17.8±1 | (15.7-19.4) | 17.8±1.1 | (15.7-19.4) | 17.8±1 | (16.2-19.4) | 17.6±1.3 | (15.3-19.5) | 17.3±1.4 | (15.3-19.5) | 18.3±1.1 | (17.2-19.3) |
| DHL | 15.3 | 14.6±0.9 | (12.6-16.3) | 14.2±0.6 | (12.6-15.6) | 14.1±0.8 | (12.6-15.6) | 14.3±0.5 | (13.7-15.0) | 15.4±0.8 | (14.0-16.3) | 15.2±0.9 | (14.0-16.3) | 15.8±0.2 | (15.6-16.0) |
| PreP | 53.6 | 51.5±2.3 | (42.7-55.2) | 51.9±1.8 | (48-55.2) | 51.8±1.8 | (49.3-55.2) | 52.1±1.9 | (48.0-54.5) | 50.5±3.1 | (42.7-53.0) | 50.1±3.8 | (42.7-53.0) | 51.4±0.7 | (50.6-51.9) |
| PreA | 77.2 | 78.3±1.5 | (75.2-81.8) | 78.5±1.4 | (75.2-81.1) | 78.8±1.3 | (76.5-81.1) | 78.1±1.5 | (75.2-80.3) | 77.8±1.8 | (76.1-81.8) | 78.2±2.0 | (76.1-81.8) | 76.8±0.3 | (76.5-77.0) |
| PreD | 45 | 47.8±2 | (39.9-50.3) | 48±1.5 | (45.5-50.3) | 48.3±1.5 | (45.5-50.3) | 47.6±1.7 | (45.6-50) | 47.3±3 | (39.9-50.2) | 47.1±3.6 | (39.9-50.2) | 47.8±1.6 | (46.3-49.5) |
| BD | 27.6 | 26.5±2.7 | (19.9-31.8) | 27.5±1.9 | (24.5-31.8) | 28.2±2.0 | (25.2-31.8) | 26.5±1.4 | (24.5-28.8) | 24.2±2.9 | (19.9-27.0) | 24.8±2.8 | (19.9-27.0) | 23.1±3.3 | (20.5-26.8) |
| PostD | 43.7 | 42.6±2 | (34.7-46) | 42.7±1.3 | (38.3-43.9) | 42.3±1.5 | (38.3-43.9) | 43.2±0.5 | (42.1-43.8) | 42.3±3.3 | (34.7-46) | 42.3±4.1 | (34.7-46) | 42.5±0.9 | (41.4-43) |
| TL | 118 | 120.5±4.2 | (100-124.2) | 121±1.3 | (118.7-123.4) | 120.6±1.5 | (118.7-123.4) | 121.5±0.7 | (120.8-122.8) | 119.2±7.5 | (100-124.2) | 118.6±9.3 | (100-124.2) | 120.5±1.7 | (118.9-122.4) |
| HL (mm) | 58.6 | 54.2±10.6 | (31.1-73.1) | 58.3±8.6 | (41.9-73.1) | 60.2±6.8 | (53-73.1) | 55.7±10.4 | (41.9-72.2) | 44.6±8.7 | (31.1-58.3) | 49.5±4.7 | (45.7-58.3) | 34.9±6 | (31.1-41.8) |
| Percentage of head length | |||||||||||||||
| SN | 22.2 | 23.1 ± 1.6 | (20.1-26.4) | 23.5 ± 1.5 | (20.9-26.4) | 23.7 ± 1.5 | (21.4-26.4) | 23.2 ± 1.5 | (20.9-25.3) | 22.4 ± 1.6 | (20.1-25) | 22.8 ± 1.7 | (20.3-25) | 21.7 ± 1.4 | (20.1-22.7) |
| ED | 22.1 | 22.5 ± 2 | (19.5-27.2) | 21.7 ± 1.5 | (19.5-26.1) | 21.3 ± 1.1 | (19.5-22.9) | 22.3 ± 2 | (19.8-26.1) | 24.3 ± 1.7 | (21.8-27.2) | 23.3 ± 1.1 | (21.8-24.9) | 26.1 ± 1 | (25.4-27.2) |
| EC | 28.9 | 27.4 ± 1.7 | (24.5-31.5) | 26.7 ± 1.2 | (24.5-29) | 26.6 ± 1 | (24.9-28.3) | 26.9 ± 1.4 | (24.5-29) | 29 ± 1.9 | (26.4-31.5) | 28.1 ± 1.7 | (26.4-30.6) | 30.6 ± 0.8 | (29.9-31.5) |
| Morphological characters | C. alpinus | Both lakes | Lake Thun | Lake Brienz | |||||||||||
| Lectotype | Non-types both sexes | Non-types | Non-types | ||||||||||||
| N-total = 30 | N-total = 21 | N-females = 12 | N-males = 9 | N-total = 9 | N-females = 6 | N-males = 3 | |||||||||
| Mean ± StDev | Range | Mean ± StDev | Range | Mean ± StDev | Range | Mean ± StDev | Range | Mean ± StDev | Range | Mean ± StDev | Range | Mean ± StDev | Range | ||
| EH | 23.7 | 22.2 ± 1.8 | (19.1-27.1) | 21.4 ± 1.4 | (19.1-23.6) | 21.1 ± 1.2 | (19.1-23) | 21.9 ± 1.6 | (19.6-23.6) | 23.9 ± 1.5 | (22.4-27.1) | 23.3 ± 0.9 | (22.4-24.6) | 25.1 ± 2 | (23.2-27.1) |
| ES | 5.7 | 5.0 ± 0.8 | (3.3-6.3) | 5.1 ± 0.8 | (3.4-6.3) | 5.0 ± 0.7 | (3.7-6.0) | 5.2 ± 0.9 | (3.4-6.3) | 4.7 ± 0.8 | (3.3-5.8) | 4.6 ± 0.9 | (3.3-5.5) | 4.9 ± 0.8 | (4.3-5.8) |
| PostO | 51.2 | 52.4 ± 1.5 | (48.9-55.4) | 52.6 ± 1.2 | (50.2-55.4) | 52.5 ± 1.3 | (50.8-55.4) | 52.8 ± 1.2 | (50.2-54.4) | 51.7 ± 1.9 | (48.9-54.8) | 52.2 ± 1.5 | (50.7-54.8) | 50.8 ± 2.6 | (48.9-53.8) |
| HD | 71.7 | 71 ± 3.7 | (65.5-79.6) | 71.6 ± 4 | (65.6-79.6) | 71.5 ± 4.6 | (65.6-79.6) | 71.8 ± 3.4 | (67.9-76.7) | 69.4 ± 2.5 | (65.5-73.2) | 69.7 ± 2.7 | (65.5-73.2) | 68.7 ± 2.3 | (66.3-70.8) |
| MW | 8.9 | 9.4 ± 0.5 | (8.4-10.4) | 9.4 ± 0.5 | (8.4-10.4) | 9.3 ± 0.6 | (8.4-10.4) | 9.4 ± 0.6 | (8.8-10.2) | 9.5 ± 0.5 | (8.7-10.2) | 9.6 ± 0.5 | (9.2-10.2) | 9.3 ± 0.5 | (8.7-9.7) |
| UJ | 27.8 | 27.4 ± 1.5 | (24.3-30.1) | 27.7 ± 1.5 | (24.3-30.1) | 27.2 ± 1.5 | (24.3-29.4) | 28.4 ± 1.3 | (26.5-30.1) | 26.8 ± 1.3 | (25.4-29.1) | 26.5 ± 1 | (25.6-28.2) | 27.3 ± 1.8 | (25.4-29.1) |
| LJ | 38.2 | 38.4 ± 1.7 | (33.8-41.4) | 38.6 ± 1.7 | (36.6-41.4) | 38.5 ± 1.8 | (36.6-41.4) | 38.7 ± 1.6 | (37-41.2) | 38.2 ± 1.8 | (33.8-39.4) | 38 ± 2.1 | (33.8-39.2) | 38.5 ± 1.3 | (36.9-39.4) |
| M | 22.5 | 20 ± 1.2 | (16.6-22.5) | 20 ± 1.1 | (17.7-22.1) | 19.7 ± 0.9 | (18.4-21) | 20.3 ± 1.3 | (17.7-22.1) | 20 ± 1.6 | (16.6-22.5) | 19.6 ± 1.6 | (16.6-21.1) | 20.8 ± 1.5 | (19.5-22.5) |
| SD | 9.9 | 10.1 ± 1.4 | (7.2-12.9) | 10.5 ± 1.3 | (8.5-12.9) | 10.5 ± 1.4 | (8.5-12.9) | 10.6 ± 1.2 | (8.7-12.2) | 9.2 ± 1.1 | (7.2-10.5) | 9.2 ± 0.8 | (7.9-10) | 9.3 ± 1.9 | (7.2-10.5) |
| SW | 13.8 | 15.6 ± 1.1 | (13.7-17.6) | 15.6 ± 1.2 | (13.7-17.6) | 15.7 ± 1.1 | (14.1-17.6) | 15.5 ± 1.4 | (13.7-17.2) | 15.7 ± 0.9 | (14.6-17.6) | 15.6 ± 0.8 | (14.6-16.7) | 16 ± 1.3 | (15.3-17.6) |
| HW | 47.1 | 50 ± 4.5 | (39.2-59.5) | 51.3 ± 4.1 | (44.2-59.5) | 51.5 ± 5 | (44.2-59.5) | 51 ± 2.8 | (46.3-55.9) | 46.9 ± 4 | (39.2-52.3) | 47.4 ± 5 | (39.2-52.3) | 45.8 ± 0.4 | (45.4-46.1) |
| IOW | 24.5 | 27.7 ± 2.2 | (22.4-32.5) | 28.3 ± 2.3 | (22.4-32.5) | 28 ± 2.4 | (22.4-32.4) | 28.7 ± 2.3 | (24.9-32.5) | 26.5 ± 1.2 | (24.9-28) | 26.5 ± 1.4 | (24.9-28) | 26.6 ± 1 | (25.5-27.4) |
| INW | 10.7 | 11.7 ± 1 | (9.5-14.1) | 12.1 ± 1 | (10.5-14.1) | 12 ± 1.1 | (10.5-14.1) | 12.1 ± 0.9 | (10.7-14) | 11 ± 0.7 | (9.5-11.9) | 10.9 ± 0.9 | (9.5-11.9) | 11.2 ± 0.4 | (10.9-11.6) |
| LJW | 10.3 | 11.1 ± 2.2 | (7.3-15.7) | 12.1 ± 1.7 | (10.1-15.7) | 11.9 ± 1.8 | (10.1-15.4) | 12.4 ± 1.6 | (10.5-15.7) | 8.8 ± 1.2 | (7.3-10.6) | 9 ± 1.2 | (7.5-10.6) | 8.5 ± 1.4 | (7.3-10.1) |
| UJW | 19.8 | 23.2 ± 2.2 | (18.4-27.2) | 23.5 ± 2.2 | (19.9-27.2) | 23 ± 2.5 | (19.9-27.2) | 24.1 ± 1.6 | (21.9-26.5) | 22.8 ± 2.2 | (18.4-25.6) | 23.7 ± 1.6 | (21.4-25.6) | 20.9 ± 2.2 | (18.4-22.8) |
| MGR | 10.6 | 10.9 ± 1.4 | (8.3-15.2) | 11.3 ± 1.4 | (9.3-15.2) | 11.3 ± 1.4 | (9.3-15.2) | 11.3 ± 1.4 | (9.5-13.2) | 9.8 ± 1 | (8.3-11.2) | 9.9 ± 1.1 | (8.3-11.2) | 9.7 ± 0.8 | (8.7-10.2) |
| LGR | 11.5 | 11.9 ± 1.2 | (10-15.2) | 12.3 ± 1.1 | (10.6-15.2) | 12.2 ± 1.2 | (10.6-15.2) | 12.5 ± 1 | (11.3-14.1) | 10.8 ± 0.7 | (10-12.3) | 10.9 ± 0.8 | (10-12.3) | 10.6 ± 0.6 | (10-11.2) |
| UA | na | 18.5 ± 1.4 | (15.6-21.5) | 18.4 ± 1.3 | (15.6-20.7) | 18.6 ± 1.5 | (15.6-20.7) | 18.2 ± 1.1 | (16.1-19.7) | 18.7 ± 1.7 | (16.4-21.5) | 18.2 ± 1.4 | (16.4-20) | 19.8 ± 1.7 | (18-21.5) |
| LA | 35 | 33.9 ± 2.1 | (28.6-38.8) | 33.5 ± 1.8 | (28.6-36.3) | 34.1 ± 1.5 | (30.9-36.3) | 32.7 ± 1.9 | (28.6-35) | 34.7 ± 2.7 | (30.4-38.8) | 34.5 ± 2.7 | (30.4-38.2) | 35.1 ± 3.2 | (32.8-38.8) |
| Meristic characters | Mode | Range | Mode | Range | Mode | Range | Mode | Range | Mode | Range | Mode | Range | Mode | Range | |
| PelvF unbranched | 1 | 1 | (1-1) | 1 | (1-1) | 1 | (1-1) | 1 | (1-1) | 1 | (1-1) | 1 | (1-1) | 1 | (1-1) |
| PelvF branched | 11 | 11 | (10-11) | 11 | (10-11) | 11 | (10-11) | 11 | (10-11) | 10 | (10-11) | 10 | (10-11) | 11 | (10-11) |
| PecF unbranched | 1 | 1 | (1-1) | 1 | (1-1) | 1 | (1-1) | 1 | (1-1) | 1 | (1-1) | 1 | (1-1) | 1 | (1-1) |
| PecF branched | 16 | 15 | (14-17) | 15 | (14-17) | 15 | (14-17) | 16 | (14-16) | 15 | (15-17) | 15 | (15-16) | na | (15-17) |
| DF unbranched | 3 | 4 | (3-4) | 4 | (3-4) | 4 | (3-4) | 4 | (3-4) | 4 | (3-4) | 4 | (3-4) | 4 | (4-4) |
| DF branched | 11 | 11 | (10-13) | 11 | (10-13) | 11 | (10-13) | 11 | (10-12) | 11 | (10-11) | 11 | (10-11) | 11 | (11-11) |
| AF unbranched | 3 | 3 | (3-4) | 3 | (3-4) | 3 | (3-3) | 3 | (3-4) | 4 | (3-4) | 4 | (3-4) | 4 | (3-4) |
| Meristic characters | C. alpinus | Both lakes | Lake Thun | Lake Brienz | |||||||||||
| Lectotype | Non-types both sexes | Non-types | Non-types | ||||||||||||
| N-total = 30 | N-total = 21 | N-females = 12 | N-males = 9 | N-total = 9 | N-females = 6 | N-males = 3 | |||||||||
| Mode | Range | Mode | Range | Mode | Range | Mode | Range | Mode | Range | Mode | Range | Mode | Range | ||
| AF branched | 12 | 12 | (10-14) | 12 | (10-14) | 12 | (10-14) | 12 | (10-13) | 11 | (11-13) | 12 | (11-13) | 11 | (11-11) |
| LS | 82 | 84 | (77-93) | 81 | (77-93) | 84 | (77-93) | 80 | (78-84) | 84 | (80-88) | 86 | (80-88) | na | (81-84) |
| PDS | 39 | 36 | (32-42) | 36 | (32-42) | 36 | (32-42) | 33 | (33-38) | 33 | (32-42) | na | (33-42) | na | (32-37) |
| TDS | 10 | 10 | (8-11) | 10 | (9-11) | 10 | (9-11) | 10 | (9-11) | 10 | (8-10) | 10 | (8-10) | 9 | (9-10) |
| TAS | 8 | 8 | (7-9) | 8 | (8-9) | 8 | (8-9) | 8 | (8-9) | 8 | (7-8) | 8 | (7-8) | 8 | (7-8) |
| TPS | 8 | 8 | (7-9) | 9 | (8-9) | 9 | (8-9) | 8 | (8-9) | 8 | (7-8) | 8 | (8-8) | 8 | (7-8) |
| UGR | 10 | 10 | (8-11) | 10 | (8-11) | 10 | (9-11) | 11 | (8-11) | 9 | (9-11) | 9 | (9-11) | na | (9-11) |
| LGR | 18 | 19 | (15-23) | 19 | (16-23) | 19 | (16-21) | 20 | (17-23) | 18 | (15-21) | 17 | (17-21) | na | (15-19) |
| total GR | 28 | 29 | (25-34) | 30 | (25-34) | 28 | (26-32) | 30 | (25-34) | 28 | (26-30) | 29 | (27-30) | 28 | (26-28) |