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. 2020 Oct 15;10(5):217–230.

Table 3.

Different methods for generation of VST

I. In vitro stimulation and expansion of VST [80,86,95,96]

• Repeated stimulation of donor derived peripheral blood mononuclear cells with antigen presenting cells (APC) pulsed with target antigens. Ultimately expansion of T cells using IL-2.
• Various antigen presenting cells include dendritic cells, monocytes, PHA blasts, B cells and artificial APC can be utilized.
• Antigen sources are comprised of whole virus/viral lysate, whole proteins, viral vectors and peptide/peptide mixtures.
Advantage and disadvantage:
    ◦ It is the first and most commonly used protocol.
    ◦ Need relatively a small volume of blood.
    ◦ Final product containing polyclonal T cells.
    ◦ This method require a long production time.

II. Direct selection of VST [80,86,95,96]

• Donor white blood cells are isolated in vitro via three different method:
    - Peptide-HLA multimers.
    - Cytokine-capture method after exposure to viral antigen.
    - Based on expression and upregulation of activation molecules on the cell surface such as CD137 (4-1BB) and CD154 (CD40L).
Advantage and disadvantage:
    ◦ It is a more rapid way to generate VST.
    ◦ This technique is basically limited to donors with specific immunological memory T cells for the virus.
    ◦ Large volume of donor blood (100-500 mL) is typically required to reach valuable cell doses.
    ◦ In vitro activation or expansion lead to exhaustion of T cells.

III. Multivirus-specific T cells [80,86,95,96]

• Generation of VST for each single virus need a separate manufacturing process and it is time and cost consuming. Different practices for the generation of multivirus-specific T cells in one single step have been established.
• This product can be used for patients with multiple refractory viral infections.
Advantage and disadvantage:
    ◦ Targeting of various viruses in a solitary product.
    ◦ Challenges comprise production time, cost, and labor.
    ◦ A potential difficulty is that the most immunodominant antigens will prevent other T cell expansions and diminish the final product of clonal diversity.

IV. Third Party VST [80,86,95]

• Manufacturing of donor-derived VST is not always conceivable because of virus seronegative donors or patients undergoing umbilical cord blood (UCB) transplantation. Third-party VST can be utilize for these patients.
• Third-party VST represent an alternative option in patients who don’t respond to donor-derived VST because of mutations in viral genome that cannot be recognized by donor T cells.
• This product derived from unmatched donors.
Advantage and disadvantage:
    ◦ Since it is not from autologous or HLA-matched sources, may increase the risk of GVHD.
    ◦ It is a partially matched product that could be recognized by the host immune system leading to rejection of VST.
    ◦ It is a products that could be immediately available in the patients with refractory infections.
    ◦ It is for “off the shelf” administration that lead to avoid any risky delay in the management of viral disease.