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. 2020 Nov 18;8:e10409. doi: 10.7717/peerj.10409

Table 1. Ethogram for the classification and definition of the behavioural elements of the observed wild boar.

Context Definition
Locomotion (L)
Walking Slow movement (pace), every leg is moved at least one step (also backwards possible).
Running Fast movement (trot and faster).
Jumping Jump over an obstacle or ditch.
Flight Abrupt escape from recent whereabouts (optionally just a few steps).
Olfactory behaviour (OB)
Sniffing Sniffing on the ground or between plants of the ground and herb layer.
Winding Sniffing in the air or at something (e.g., rubbing tree, camera).
Defecating Emptying of the gut.
Urinating Total drain of the bladder.
Vigilance behaviour (VB)
Getting frightened Short wince of the whole body.
Pausing Freeze of motion with alert view and potential additional head lift and look about.
Laying down Young boar presses its body abrupt even on the ground.
Guarding Alert milling around, with lifted head and tail, obvious tense posture.
Foraging behaviour (FB)
Pawing Pawing in the ground (e.g., soil, leaves) with a foreleg.
Rooting Rooting in the ground (e.g., soil, leaves) with the snout, also with brushing big branches aside.
Salt ingestion Ingestion of salt at a salt lick by licking, nibbling.
Sucking attempt Young boar attempt to suck on the sow’s teats or briefly suck at the standing sow.
Suckling Young boar are suckled by the lying sow.
Chewing Uniform opening and closing of the mouth after foraging (feeding not visible).
Feeding attempt Young boar takes soil/stone into its mouth.
Feeding Ingestion of food with the mouth and chewing afterwards.
Drinking Ingestion of water with the mouth.
Comfort behaviour (CB) –personal hygiene behaviour
Stretching Increasing the distance of the hind legs to the forelegs and slightly spreading of the hind legs while simultaneously scuttling with the forelegs.
Shaking Moving its body strongly, briefly and fast back and forth while standing.
Rubbing Rubbing one’s body against a tree or another wild boar.
Nibbling Nibbling/rubbing of the open mouth against the rubbing tree.
Scratching Scratching one’s body with the hind legs.
Scratching one’s bottom Rubbing one’s bottom against the ground while sitting.
Rolling Rubbing one’s body against the ground.
Wallowing Laying down (and optionally wallowing) in muddy water.
Social interaction (SI)
Active socio negative interaction
Threating Keeping another wild boar at distance by threating behaviour.
Pushing away softly Pushing another wild boar softly away with the head, the side of the body or the bottom.
Chasing away A wild boar runs after another wild boar, which departs itself afterwards.
Snout knock A wild boar knocks its head bottom-up in the direction of another wild boar (with/without touching).
Passive socio negative interaction
Retreating A wild boar increases the distance to another wild boar, which emitted socio negative behaviour before.
Socio positive interaction
Nose-to-nose contact Sniffing at or touching the snout region (being sent of one or both wild boar, also at distance).
Nose-to-body contact A wild boar sniffs at or touches another wild boar with the snout at its body or legs.
Playing Playful behaviour against other wild boar (e.g., exercise fights, apparent copulation attempt).
Sexual behaviour (SB)
Copulation attempt A wild boar climbs the bottom of another wild boar.