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. Author manuscript; available in PMC: 2021 May 28.
Published in final edited form as: Adv Drug Deliv Rev. 2020 May 28;154-155:245–273. doi: 10.1016/j.addr.2020.05.006

Table 8.

Various liposomal active targeting ligands for active components of CHMs targeted to cancer cells and the TME

Targeting ligands Targeting site Entrapped components Drug loading Cancer model Main results Reference
Over-expressed receptors on the surface of cancer cells
β-Galactose Asialoglycoprotein receptors Glycyrrhetinic acid (GA) Film dispersion method In vivo
Kunming mice
In vitro
HepG2 cells
In vivo
Galactosylated GA-liposomes (Gal-GA-LP) showed long-circulating profile in Kunming mice with the MRT of Gal-GA-LP being 1.48- and 1.3-fold increasement when compared with GA solution and nontargeting GA-loaded liposomes, respectively.
In vitro
Gal-GA-LP showed 1.4-fold higher drug concentration in cells than that of GA-liposomes.
[420]
Oridonin (ORI) Ethanol injection method In vivo
Kunming mice
In vivo
The MRT of ORI-loaded liposomes modified with galactose (NOH-ORI-LP) was 5.56-fold longer than that of ORI solution in mice, with lower clearance from liver.
[421]
Lactose Matrine (MA) Reversed-phase evaporation method In vivo
Kunming mice
In vitro
HepG2 cells
In vivo
The accumulation of MA in targeting liposomes in the mice liver was 2.7 times higher than in the spleen, 3 times higher than in the lung, 6.6 times higher than in the kidney, and 8.5 times higher than in the heart.
In vitro
Inhibitory rate of targeting liposomes of MA was 1.6- and 1.83-fold higher than those of conventional MA liposomes and MA solution at 0.5 mg/mL concentration, respectively.
[422]
Transferrin (Tf) Transferrin receptors Tetrandrine and Vincristine Film dispersion method In vivo
Glioma-bearing ICR mice model
In vitro
C6 cells, C6/ADR cells and murine brain microvascular endothelial cells (BMVECs)
In vivo
Tf targeting liposomes improved accumulation in brain tumor tissue with fluorescent signal maintained up to 48 h and animals exhibited prolonged survival time (42.67 ± 3.56 days) compared with the saline group.
In vitro
The targeting liposomes showed significantly prolonged circulation time and increased accumulation in tumors.
[423]
Lactoferrin (Lf) Honokiol and Daunorubicin Film dispersion plus ammonium sulfate gradient methods In vivo
Glioma-bearing ICR mice model
In vitro
C6 cells and BMVECs
In vivo
The targeting liposomes improved accumulation in brain tumor tissue indicated by fluorescence probe and conferred prolonged survival time.
In vitro
Lf modified daunorubicin plus honokiol liposomes enhanced drug transportation across the blood-brain barrier, inhibited C6 cells invasion, and destroyed vasculogenic mimicry channels.
[424]
Folic acid Folate receptors (FR) Curcumin (Cur) Film dispersion method In vitro
KB, Hela, and A549 cells
In vitro
FR-positive cells endocytosed more FR-targeting liposomes than nontargeted liposomal Cur. Targeted liposomes more effectively inhibited cellular proliferation and caused higher dose- and time-dependent apoptosis. KB cells were more sensitive to FR-targeting liposomal Cur than Hela cells.
[425]
Baicalin (Bai) Film hydration and homogenization In vitro
HeLa cells
In vitro
Cytotoxicity and cellular uptake of Bai-loaded FR-targeting liposomes were higher than that of non-targeted liposomes.
[426]
Ursolic acid (UA) Film hydration and extrusion In vivo
Balb/c nude mice xenografted oral cancer (KB cells)
In vitro
KB cells
In vivo
Compared with passive targeting liposomes, FA-PEG modified UA liposomes significantly inhibit oral tumor growth.
In vitro
The EE% of UA was more than 85%. FR targeting UA PEGylated liposomes showed improved tumor cell uptake, proliferation inhibition, and apoptotic effects as compared with nontargeting PEGylated liposomes.
[427]
Glycyrrhetinic acid (GA) GA receptors Ginsenoside Rh2 Film hydration method In vitro
SMMC-7721 cells
In vitro
IC50 of GA-modified liposomes in SMMC-7721 cells increased by 0.5- and 0.6-fold compared with Rh2 solution and Rh2-loaded liposomes, respectively.
[428]
Baicalin Film hydration and ethanol injection methods \ The size of GA modified baicalin liposomes was 150 nm with EE% at 41.8%. [429]
Brucine Ethanol injection method \ The formulation of GA modified brucine liposomes was optimized with a lipid to cholesterol (w/w) ratio of 6, lipid to drug (w/w) ratio of 15, and lipid to GA (w/w) ratio of 35 (g/g). The size of targeted liposomes was 147.2 nm and the EE% of brucine was 82.8%. [430]
Wogonin Reversed-phase evaporation method In vivo
ICR mice xenografted liver cancer (HepG2 cells)
In vitro
HepG2, L-02 and LX-2 cells
In vivo
GA modified liposomes rapidly and highly accumulated in the tumor and liver shortly after administration. The tumor inhibitory ratio of targeted liposomes was 1.7-fold higher than that of nontargeted liposomes.
In vitro
GA-modified liposomes showed 1.46-fold higher growth inhibition than that of non-targeted liposomes in HepG2 cells.
[428]
Cur Ethanol injection method In vitro
HepG-2 and H22 cells
In vivo
GA modified liposomes loaded with Cur inhibited tumor growth, reduced tumor microvascular density and regulated the expression of Caspases3 and VEGF proteins in H22 tumor tissues.
In vitro
GA modified liposomes induced more HepG2 cellular apoptosis and tumor growth inhibition.
[431]
Epigallocatechin 3-gallate (EGCG) 67LR Doxarubicin Film hydration method In vivo
C57BL/6 mice melanoma (B16F10 cells)
In vivo
Targeting liposomes showed superior anti-tumor effect without observed side effects.
[432]
Targeting cytoplasmic organelles
Mitochondrion Dequalinium (DQA) Resveratrol Film dispersion and extrusion In vivo
Balb/c nude mice xenografted resistant lung cancer (A549/cDDP cells)
In vitro
A549 cells and A549/cDDP cells
In vivo
The combination administration of targeting resveratrol liposomes with vinorelbine liposomes exhibited superior prohibition of tumor growth.
In vitro
The increased mitochondrial uptake and solubility of resveratrol in targeting liposomes promoted inhibitory cell activity by triggering cytochrome C release.
[433]
Berberine Film dispersion and extrusion In vivo
Balb/c nude mice xenografted breast cancer (MCF-7 dissociated cancer stem cells)
In vitro
MCF-7 cells
In vivo
The synergistic effect of targeting berberine liposomes and paclitaxel liposomes exhibited the strongest anti-tumor effect.
In vitro
Targeting berberine liposomes (1, 5, and 10 μM) could synergistically enhance the toxicity of paclitaxel liposomes in a dose-dependent manner, while achieving selective mitochondrial accumulation.
[434]
4-Carboxybutyl triphenylphosphonium bromide (TPP) or DQA Resveratrol Film dispersion and extrusion In vitro
B16F10 cells
In vitro
TPP and DQA conjugated with DSPE-PEG liposomes carrying resveratrol caused better cellular uptake and selective accumulation in mitochondria.
[435]
Targeting the TME
Hyaluronic acid (HA) CD44 and the receptor for hyaluronan-mediated motility Curcumin (Cur) Reversed-phase evaporation method In vitro
HepG2 and A549 cells
In vitro
The binding rate of HA-lipid to Cur-liposomes was 72%. The IC50 values were found to be 0.054, 0.032 and 0.021 mol/ml for free Cur, nontargeted Cur liposomes and modified Cur liposomes, respectively, after incubation with A549 cells overexpressing the CD44 receptor.
[436]
Matrine Film dispersion, reversed-phase evaporation and ethanol injection methods In vivo
Balb/c mice xenografted liver cancer (H22 cells)
In vivo
The inhibition rate of HA-modified MA liposomes on H22 hepatocarcinoma was similar to that of cyclophosphamide, and 2.0- and 1.4- fold higher than that of MA solution and MA conventional liposomes, respectively.
[437]
Surface modification with antibodies
Anti-CD44 antibody CD44 receptors Timosaponin AIII (TAIII) Film evaporation and ultrasonic methods In vivo
Balb/c nude mice xenografted liver cancer (HepG2 cells)
In vitro
HepG2 cells
In vivo
CD44 targeting liposomes showed significantly inhibition on tumor growth, which was 7.2- and 1.3- fold greater than that of free drug and non-targeted liposomes, respectively.
In vitro
Targeting liposomes were more toxic to tumor cells with lower IC50 than non-targeting liposomes.
[438]
Trastuzumab Human epidermal growth factor receptor-2 (HER2) Curcumin and Resveratrol Film evaporation and extrusion method In vitro
JIMT1 an MCF-7 cells
In vitro
The immunoliposomes of curcumin and resveratrol showed notably higher cytotoxicity, compared to free drugs. The finding was more prominent for curcumin than resveratrol.
[439]
Surface modification with peptides
T7 (HAIYPRH) peptide Transferrin receptors Quercetin Film hydration method In vivo
Balb/c nude mice with orthotopic lung cancer (A549-Luc cells)
In vitro
A549-Luc cells
In vivo
T7 modified liposomes significantly delayed tumor growth and enhanced the lifespan of mice following pulmonary administration.
In vitro
T7 modified liposomes showed higher cellular uptake and significantly higher tumor-spheroid growth inhibition.
[440]
NGR (Asn-Gly-Arg) CD13 Triptolide Film hydration method In vitro
Human umbilical vein endothelial cells (HUVEC)
In vitro
Triptolide-loaded liposomes modified with NGR showed stronger cellular toxicity with lowest IC50 on HUVECs at 11 μM.
[441]
Brucine Ammonium sulfate gradient method In vivo
Balb/c nude mice xenografted fibrosarcoma (HT1080 cells)
In vitro
HT1080 cells and HUVEC
In vivo
The highest fluorescence signals in tumors were found in animals treated with NGR-modified liposomes with HSPC and SPC together than that with HSPC or SPC used alone.
In vitro
The composition of lipids did not exhibit significant influence on EE% of brucine. The cellular uptake of NGR-modified liposomes was about 1.5-fold higher than that of non-targeted liposomes.
[442]
RGD (Arg-Gly-Asp) Integrins Tetrandrine and Vinorelbine Film dispersion followed by ammonium sulfate gradient method In vivo
Glioma-bearing ICR mice model (Doxorubicin-resistant C6 cells)
In vitro
C6 cells and BMVECs
In vivo
RGD modified vinorelbine and tetrandrine liposomes prolonged elimination half-life compared with free drug, and extended survival times. Strong fluorescent signals of targeting liposomes were observed in tumors within 24 h after administration, while the signals of free drugs was in the liver.
In vitro
Targeted liposomes significantly enhanced transportation across the BBB and uptake in tumor cells.
[443]
Matrine Film dispersion followed by pH-gradient method In vitro
Bcap-37, HT-29 and A375 cells
In vitro
RGD-PEG-DSPE was employed to obtain matrine liposomes with an EE% of 83.13%. Compared with free matrine, targeting liposomes induced stronger inhibition of proliferation and induced more apoptosis in tumor cells.
[444]
Shikonin (SHK) Film hydration method In vitro
MDA-MB-231 cells
In vitro
RGD modified SHK-loaded liposomes with EE% of 94.9% greatly induced cellular apoptosis by increasing the ratio of Bcl-2-associated X protein (Bax)/Bcl-2, and inhibited cellular proliferation, migration and invasion, as compared with nontargeted SHK-loaded liposomes.
[445]
Octaarginine (R8) Syndecan 4 Dihydroartemisinin and Epirubicin Film dispersion method In vitro
A549 cells
In vivo
R8 modified liposomes increased the selective drug accumulation at tumor sites and showed a targeting capable of reducing tumor volume.
In vitro
R8 modified liposomes exhibited powerful cytotoxicity on A549 cells, effectively suppressed vasculogenic mimicry channels, and regulated metastasis related proteins.
[446]
R8GD (RGD and R8) Integrins Emodin Film dispersion method In vivo
Balb/c nude mice xenografted breast cancer (MDA-MB-435S cells)
In vitro
MDA-MB-435S cells
In vivo
Strong fluorescence signals of R8GD modified liposomes were found in tumors within 24 h after administration. R8GD modified daunorubicin liposomes significantly inhibited tumor growth with the combination of R8GD modified emodin liposomes. Their efficacy was 3-fold stronger than that of nontargeting liposomes combination.
In vitro
The cellular uptake was increased for targeting liposomes.
[447]
HIV protein transactivator of transcription (TAT) together with HA Syndecan 4 Curcumin and Celecoxib Film hydration method In vivo
Balb/c mice with orthotopic breast cancer (4T1 cells)
In vitro
RAW264.7, HUVEC and 4T1 cells
In vivo
NF-κB essential modulator-binding domain (NBD) fused with TAT peptide modified liposomes co-delivering curcumin and celecoxib were coated with HA (HA/TN-CCLP). HA/TN-CCLP could prolong circulation time, increase tumor accumulation, and block lung metastasis.
In vitro
HA/TN-CCLP improved cytotoxicity, inhibited migration, and enhanced anti-inflammation effect, when compared to nontargeted liposomes.
[448]
tLyp-1 (sequence CGNKRTR) Neuropilin-1 Ginsenoside CK and Parthenolide Film hydration method In vivo
Balb/c nude mice xenografted lung cancer (A549 cells)
In vitro
A549 cells
In vivo
Targeting liposomes with hydrophilic PEG shell demonstrated stronger antitumor effect than free drugs combination without observed toxicity.
In vitro
The cellular uptake of targeted liposomes was through micropinocytosis and the liposomes were escaped from lysosome in a time-dependent manner.
[449]
Surface modification with polymers
D-alpha tocopheryl polyethylene glycol 1000 succinate (TPGS) \ Luteolin Film-dispersion method In vivo
Balb/c nude mice xenografted lung cancer (A549 cells)
In vitro
A549 cells
In vivo
TPGS coated liposomes loaded with luteolin were significantly accumulated in tumor sites. The modified liposomes exhibited higher tumor inhibition rate to 51.7% than that of free luteolin.
In vitro
TPGS coated liposomes improved cytotoxicity in A549 cells by increasing the Bax/Bcl-2 ratio.
[450]
\ Ginsenoside compound K Film hydration method In vivo
Balb/c nude mice xenografted lung cancer (A549 cells)
In vitro
A549 cells
In vivo
Targeted liposomes delivered ginsenoside compound K into tumor sites, enhanced its permeability and retained it in tumor cells.
In vitro
The EE% of targeted liposomes was 98.4%, and the size was (119.3±1.4) nm at the ratio of 7:3 (phospholipid: TPGS). The IC50 of targeted liposomes was significantly reduced to 25% of that of free drugs.
[451]