Table 1.

Network metrics used in this study. In the "Range of values" column, square brackets indicate that a range includes the endpoint and parentheses indicate that a range excludes the endpoint. In each example diagram, nodes of different colors represent different habitat types or land cover classes

Name	Definition	Ecological interpretation	Range of values	Example
Network-level metrics
Edge density	The proportion of potential connections in the network that are realized	Landscape connectedness	(0,1]
Assortativity	The tendency of nodes with similar properties to be connected to one another	Connectivity among habitats of the same vs. of different types	[-1,1]
Modularity	The ability of a network to be divided into communities, where there are few edges between communities	Aggregation of groups of patches, “functional spatial structure” [46]	[0,1]
Node-level metrics
Degree centrality	The number of links of a focal node. In a directed network, can be in-degree (the number of incoming links) or out-degree (the number of outgoing links)	The potential number of other patches that a contaminant, nutrient, etc. could directly spread to (out-degree) or come from (in-degree)	(0,N] (N=# of nodes in net-work)
Betweenness centrality	The fraction of shortest paths between nodes that pass through the focal node	Role of a patch as a “stepping stone” that connects otherwise-separated groups of patches	[0,1]
Node size	Sum of all edge weights entering and leaving a node	Number of visits to a patch	[1,Infinity)