Percentages of “Unique lifted-over to D. triauraria” represent number out of total boundaries or domains. Percentages of orthologous, non-orthologous, and subcategories of boundaries and domains represent number out of “Unique lifted-over to D. triauraria”. Truncated and expanded domains do not meet the 90% reciprocal overlap criterion due to large insertion/deletion mutations that have created asymmetry in TAD size between the two species. Missing boundaries and domains are those that could not be lifted-over to D. triauraria.