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. 2020 Dec 9;7:584162. doi: 10.3389/fvets.2020.584162
Publication/Reference number Study Birds observeda Species (breed) Days on substrateb Enclosure area (m2)c Stocking densityd Animals/m2 /daye Substrate area (m2)f Substrate depth (cm)
Jong et al., 2007 (38) 1 16 Layer 21.00 6.75 0.59 0.03 1.13–2.25 5.0
Sanotra et al., 1995 (39) 2 12 Layer 0.04 0.41 4.94 118.52 0.03–0.04 4.0
Shields et al., 2004 (40) 3 52 Broiler 4.00 2.31 0.87 0.22 0.37–0.82 10.0
Villagra et al., 2014 (41) 4 32 Broiler 0.05 4.00 2.00 41.14 1.00 15.0
Shields et al., 2005 (42) 5 60 Broiler 49.00 9.30 1.08 0.02 4.65 2.5
Toghyani et al., 2010 (43) 6 80 Broiler 42.00 5.76 3.47 0.08 1.44 2.0
van Liere et al., 1990 (44) 7 23 Layer 5.00 3.90 1.54 0.31 0.36 12.0
Hogan and Vestergaard, 1992 (45) 8 10 Jungle Fowl 0.02 0.48 4.17 171.43 0.12 2.0
Petherick and Duncan, 1989 (46) 9 48 Layer 0.02 5.76 1.04 50.00 0.13 4.0
Guinebretière et al., 2014 (33) 10 60 Layer 4.00 1.05 3.81 0.95 0.49 3.0
a

Total number of birds observed in each study (some studies picked focal birds).

b

Number of days birds were allowed access to tested substrates (in enclosure area). Small numbers represent studies that moved animals into a separate area to observe.

c

The size of the total enclosure, encompassing all substrate areas, in m2.

d

Animals per m2; Stocking density = animals per group (not shown; some studies used focal birds) ÷ enclosure area (m2).

e

Proxy for degree of substrate soiling (higher numbers indicate more soiling); animals/m2/day = stocking density ÷ days on substrate.

f

The area (m2) occupied by one substrate within the enclosure.