Effects of olfactory and/or mating stimulation on plasma luteinizing hormone (LH) and testosterone levels in intact male rats. The changes in the mean plasma LH levels of intact male rats in each group that were exposed to the clean bedding, the female-soiled bedding, or female-soiled bedding and mated with an ovariectomized (OVX) + estradiol-17β (E2) rat (A). The changes in the mean plasma testosterone levels in each group (B). Arrows indicate the onset of each bedding exposure and/or mating with a female. †, values indicating significant difference compared with the value in the males exposed to clean bedding and female-soiled bedding (P < 0.05, two-way ANOVA). *, values indicating significant difference compared with the value in the males exposed to clean bedding (P < 0.05, two-way ANOVA).