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. 2021 Jan 1;8:7. doi: 10.1038/s41438-020-00437-3

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© The Author(s) 2020

Open Access This article is licensed under a Creative Commons Attribution 4.0 International License, which permits use, sharing, adaptation, distribution and reproduction in any medium or format, as long as you give appropriate credit to the original author(s) and the source, provide a link to the Creative Commons license, and indicate if changes were made. The images or other third party material in this article are included in the article’s Creative Commons license, unless indicated otherwise in a credit line to the material. If material is not included in the article’s Creative Commons license and your intended use is not permitted by statutory regulation or exceeds the permitted use, you will need to obtain permission directly from the copyright holder. To view a copy of this license, visit http://creativecommons.org/licenses/by/4.0/.

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Fig. 8 — a OpWRKY2 positively regulates camptothecin biosynthesis by binding a W-box in the OpTDC promoter and activating the expression of OpTDC. OpWRKY1 negatively regulates camptothecin biosynthesis by binding two W-boxes in the OpCPR promoter, while OpWRKY3 positively regulates camptothecin biosynthesis by binding one W-box in the OpCPR promoter. b Camptothecin biosynthetic pathway in O. pumila. Tryptophan decarboxylase (OpTDC) catalyzes the conversion of tryptophan to tryptamine. Dashed lines represent interactions that may be indirect.