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. 2021 Jan 4;22:4. doi: 10.1186/s13059-020-02222-w

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© The Author(s) 2021

Open AccessThis article is licensed under a Creative Commons Attribution 4.0 International License, which permits use, sharing, adaptation, distribution and reproduction in any medium or format, as long as you give appropriate credit to the original author(s) and the source, provide a link to the Creative Commons licence, and indicate if changes were made. The images or other third party material in this article are included in the article's Creative Commons licence, unless indicated otherwise in a credit line to the material. If material is not included in the article's Creative Commons licence and your intended use is not permitted by statutory regulation or exceeds the permitted use, you will need to obtain permission directly from the copyright holder. To view a copy of this licence, visit http://creativecommons.org/licenses/by/4.0/. The Creative Commons Public Domain Dedication waiver (http://creativecommons.org/publicdomain/zero/1.0/) applies to the data made available in this article, unless otherwise stated in a credit line to the data.

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Fig. 1 — Study design and sample sets used in this study. Forty patients (sample set 1) were enrolled in this study. Genomic and transcriptomic profiling were performed to specimens from 17 patients of sample set 1. Experimental validation was performed using specimens from 35 patients of sample set 1, 105 non-metastatic patients from sample set 2, and two human PDAC cell lines from sample set 3. To explore the biological and clinical significance of identified events, molecular and survival data from previously published non-metastatic PDACs were used (sample set 4)