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Three novel genetic risk loci reaching genome-wide significance. (A) rs6427389 at 1q23.1 (FCRL3); (B) rs6942325 at 6p25.3 (DUSP22, IRF4); and (C) rs2240335 at 1p36.13 (PADI4). Shown are P values obtained in GWAS discovery, in the combined analysis of GWAS and Chinese validation samples, and/or in silico European samples (fixed-effects meta-analysis). Chr1, chromosome 1; chr6, chromosome 6; val, validation.
