Table 2.

Simulation of mating nucleus colonies in BEEHAVE.

Clothianidin	Brood survival experiments	Brood survival model	ProteinNurses Modifier_Exposed
control	0.8	0.92 ± 0.14	1
1 µg/L	0.4	0.38 ± 0.16	0.82
10 µg/L	0.2	0.20 ± 0.12	0.77
100 µg/L	0.0	0.02 ± 0.03	0

Modification of the factor ProteinNursesModifier_Exposed in BEEHAVE simulations resulted in brood survival rates similar to experimental data obtained in field experiments. The factor ProteinNursesModifier_Exposed reduces the protein content of the jelly fed to the larvae (ProteinFactorNurses) in the model. A wide range of values for ProteinNursesModifier_Exposed was tested, and the best fits are shown based on a comparison of modeled and experimental brood survival.