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. 2021 Jan 11;11:380. doi: 10.1038/s41598-020-79654-8

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© The Author(s) 2021

Open Access This article is licensed under a Creative Commons Attribution 4.0 International License, which permits use, sharing, adaptation, distribution and reproduction in any medium or format, as long as you give appropriate credit to the original author(s) and the source, provide a link to the Creative Commons licence, and indicate if changes were made. The images or other third party material in this article are included in the article's Creative Commons licence, unless indicated otherwise in a credit line to the material. If material is not included in the article's Creative Commons licence and your intended use is not permitted by statutory regulation or exceeds the permitted use, you will need to obtain permission directly from the copyright holder. To view a copy of this licence, visit http://creativecommons.org/licenses/by/4.0/.

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Schematic presentation of different nucleosome/chromatosome phases. (A). Isotropic solution with repulsive interaction between the particles. (B). Columnar phase without inter-columnar order. The stacking distance, h, characterises the separation between nucleosomes or chromatosomes within the column. (C). The lamella-columnar phase observed for the 177 bp chromatosome; a_H is an inter-columnar distance within a bilayer; d_L is the distance between bilayers. The yellow dots indicate the facing direction of the nucleosome dyad or location of linker DNA in the chromatosome. Adapted from⁶¹ with permission from Elsevier.