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. 2021 Jan 25;12:574. doi: 10.1038/s41467-020-20887-6

Table 2.

Mutation frequency driver genes.

Metastatic CRC Primary CRC—(TCGA, combined studies cBioportal) Metastatic CRC—(Yaeger et al.)
Gene dN/dS q-value Mutations (N) Mutations (%) Mutations (N) Mutations (%) Fisher p-value FDR Hochberg % change in meta Mutations (N) Mutations (%) Fisher p-value FDR Hochberg
TP53 0 317 73.9 1123 57.6 2.04E-10 4.7E-09 16.3 246 76.6 0.395 1
ZFP36L2 0 42 9.8 97 5.0 3.61E-04 0.008 4.8 Not present
KRAS 0 203 47.3 744 38.2 5.88E-04 0.012 9.1 127 39.6 0.037 0.487
APC 0 336 78.3 1372 70.4 8.86E-04 0.018 7.9 241 75.1 0.335 1
PIK3CA 0 68 15.9 445 22.8 0.001 0.023 −7.0 49 15.3 0.840 1
B2M 5.37E-03 8 1.9 91 4.7 0.007 0.128 −2.8 2 0.6 0.202 1
SMAD4 0 74 17.2 243 12.5 0.010 0.164 4.8 47 14.6 0.367 1
ATM 9.50E-04 33 7.7 227 11.6 0.017 0.266 −4.0 18 5.6 0.306 1
FBXW7 0 51 11.9 301 15.4 0.061 0.912 −3.6 25 7.8 0.068 0.814
AMER1 0 37 8.6 209 10.7 0.220 0.913 −2.1 11 3.4 0.004 0.067
ARID1A 1.13E-09 39 9.1 201 10.3 0.480 0.913 −1.2 15 4.7 0.022 0.333
BCL9 5.27E-02 28 6.5 107 5.5 0.419 0.913 1.0 Not present
BCL9L 2.24E-06 27 6.3 133 6.8 0.750 0.913 −0.5 Not present
BRAF 0 56 13.1 273 14.0 0.644 0.913 −1.0 38 11.8 0.657 1
ELF3 5.37E-03 7 1.6 51 2.6 0.299 0.913 −1.0 Not present
LMTK3 1.33E-02 15 3.5 56 2.9 0.530 0.913 0.6 not present
NRAS 0 26 6.1 125 6.4 0.913 0.913 −0.4 14 4.4 0.329 1
PTEN 4.30E-08 17 4.0 123 6.3 0.069 0.913 −2.3 14 4.4 0.854 1
RNF43 2.20E-02 28 6.5 162 8.3 0.239 0.913 −1.8 21 6.5 1.000 1
SMAD3 5.68E-03 11 2.6 72 3.7 0.309 0.913 −1.1 11 3.4 0.518 1
SOX9 0 41 9.6 177 9.1 0.782 0.913 0.5 16 5.0 0.025 0.352
TCF7L2 1.07E-09 50 11.7 177 9.1 0.103 0.913 2.6 19 5.9 0.007 0.116
TGIF1 1.33E-02 18 4.2 62 3.2 0.300 0.913 1.0 Not present

Twenty-three genes identified as putative driver genes using the ratio of nonsynonymous to synonymous substitutions caused by the somatic nucleotide mutations (SNV and InDels; dN/dS analysis). P-values are derived from the Fisher exact test (two-sided) and corrected for multiple testing using the FDR (Hochberg) method.