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. 2021 Jan 28;11:2595. doi: 10.1038/s41598-021-81587-9

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© The Author(s) 2021

Open Access This article is licensed under a Creative Commons Attribution 4.0 International License, which permits use, sharing, adaptation, distribution and reproduction in any medium or format, as long as you give appropriate credit to the original author(s) and the source, provide a link to the Creative Commons licence, and indicate if changes were made. The images or other third party material in this article are included in the article's Creative Commons licence, unless indicated otherwise in a credit line to the material. If material is not included in the article's Creative Commons licence and your intended use is not permitted by statutory regulation or exceeds the permitted use, you will need to obtain permission directly from the copyright holder. To view a copy of this licence, visit http://creativecommons.org/licenses/by/4.0/.

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Niigata prefecture map with the sites where adult stages of Ixodes spp. were collected (Circles), and the occurrence of SFGR (filled circle: SFGR Positive sites, open circle: SFGR Negative sites). From the left, “A” corresponds to I. ovatus and Rickettsia asiatica sites; “B” corresponds to I. monospinosus and R. helvetica sites, and “C” are the I. nipponensis and R. monacensis sites. Maps created using the Geographical Survey Institute Map Vector from the Geospatial Information Authority of Japan (https://maps.gsi.go.jp/).