Effects of storage temperature at 4°C (■), -18°C (▴), and -80°C (●) on normal germination in S. koreensis Andersson. Seeds were stored for up 52 months with water contents controlled to 3% (A), 6% (B), 9% (C), 12% (D), 18% (E), or 24% (F) (fresh weight basis). Storage temperature (ST)***, seed water content (SWC)***, storage period (SP)***, ST × SWC***, ST × SP***, SWC × SP***, ST × SWC × SP***. All values are means of four replicates ± SD (n=4). *** indicates significant difference at p < 0.001.