Table 3. Summary of clock model operators introduced throughout this article.

Operator	Description	Parameters
AdaptiveOperatorSampler	Samples sub-operators proportionally to their weights, which are learned (see Adaptive operator weighting).	${\overrightarrow{\mathcal{R}}}^{},\sigma ,t\mathcal{T}$
SampleFromPrior	Resamples a random number of elements from their prior (see Adaptive operator weighting).	${\overrightarrow{\mathcal{R}}}^{},\sigma$
NarrowExchangeRate	Moves a branch and recomputes branch rates so that their genetic distances are constant (see Narrow exchange rate).	${\overrightarrow{\mathcal{R}}}^{},\mathcal{T}$
LeafAVMVN	Proposes new rates for all leaves in one move (see An adaptive leaf rate operator) [30].	${\overrightarrow{\mathcal{R}}}^{}$

Pre-existing clock model operators are summarised in Table 1. t denotes node heights while $\mathcal{T}$ denotes the whole tree, including topology.