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(A) Micromanipulation force measurement schematic of how the technique works. A force pipette (fp) and a pull pipette (pp) are used to extend the nucleus (nuc) (B) A single isolated nucleus can be measured before and after biochemical treatment, providing fine-tuned control over nuclear chromatin mechanical and imaging measurements. The nuclear spring constant (knuc) increases during chromatin condensation and decreases after chromatin digestion.
