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. 2021 Feb 25;11:4586. doi: 10.1038/s41598-021-84070-7

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© The Author(s) 2021

Open Access This article is licensed under a Creative Commons Attribution 4.0 International License, which permits use, sharing, adaptation, distribution and reproduction in any medium or format, as long as you give appropriate credit to the original author(s) and the source, provide a link to the Creative Commons licence, and indicate if changes were made. The images or other third party material in this article are included in the article's Creative Commons licence, unless indicated otherwise in a credit line to the material. If material is not included in the article's Creative Commons licence and your intended use is not permitted by statutory regulation or exceeds the permitted use, you will need to obtain permission directly from the copyright holder. To view a copy of this licence, visit http://creativecommons.org/licenses/by/4.0/.

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Significant associations—(A): BALF5, (B): BBRF1, (C): BRLF1:p.Lys316Glu. The x-axis represents the chromosomal position and the y-axis displays the -log10(p-value). Colour alternates between chromosomes. Regions that contain statistically significant SNP are presented in red (top SNP ± 400 Kbp). The light grey dashed line represents the GWAS significance threshold of 5 × 10^–8, the dark grey dashed line the G2G threshold of 1.09 × 10^–10. This figure was produced using R⁶⁵.