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. 2021 Feb 24;10:1–27. doi: 10.2147/OV.S268426

Table 3.

List of Armed oHSVs for GBM Treatment

Armed Transgene oHSV Strain Genomic Modification GBM Model PFU (Doses) Results and References
Cytokines G47∆-mIL12 F ∆ICP6, -/-γ34.5, ∆ICP47, +LacZ, +mIL-12 005 GSC 5×105 (2 doses) G47∆-mIL12 significantly extended survival with 10% mice surviving long term compared to 0% in control virus (G47∆-Empty) group.33
M002 F -/-γ34.5, +mIL-12 Patient’s GBM tumors 1×107 (1 dose) - Pediatric GBM samples with high nectin-1 expression: M002 treatment led to tumor eradication in 8/10 animals compared to 3/10 animals in the G207 group.58
- Pediatric GBM samples with low nectin-1 expression and adult GBM samples: M002 treatment showed trend of improved median survival versus G207 treatment.58
oHSV-Flt3-L F ∆ICP6, -/-γ34.5, ∆ICP47, +LacZ, +Flt3-L CT-2A 2x106 (1 dose) Treatment with G47Δ-Flt3L led to 40% long-term survivors compared to 0% in G47Δ-Empty treatment.60
oHSV-TRAIL F ∆ICP6, -/-γ34.5, ∆ICP47, +LacZ, +TRAIL LN229-FmC GBM 2×108 (2 doses) The median survival of mice treated with oHSV-TRAIL was 69 days, which was significantly longer than PBS group (50.5 days) or unarmed oHSV group (49 days).61
GSC23-FmC and GSC31 GSC23-FmC - 2×106 (2 doses) and GSC31 2×106 (1 dose) - GSC23-FmC: The median survival of mock group (44 days) was significantly lower than oHSV-TRAIL (55 days).62
- GSC31: oHSV-TRAIL significantly extended median survival (81 days) with 40% long-term survivors compared to 0% (median survival - 26 days) in the mock mock group.62
Angiogenic inhibitors G47∆-mAngio F ∆ICP6, -/-γ34.5, ∆ICP47, +LacZ, +mAngio U87 5×106 (2 doses) - Subcutaneous U87 model: G47∆-mAngio delayed tumor growth by 4 days compared to G47∆-E.67
- Intracranial U87 model: Treatment with G47∆-mAngio resulted in 40% of mice surviving over 150 days and significantly prolonged animal survival compared to G47∆-E.67
1×106 (1 dose) The median survival of G47Δ-mAngio was 66 days, which was significantly longer than G47Δ-Empty-treated group (53 days).57
MGG4 1×106 (1 dose) The median survival of G47Δ-mAngio was 113 days, which was significantly longer than G47Δ-Empty group (98 days).57
VAE F ∆ICP6, -/-γ34.5, +endo-angio Human stable GSC lines Subcutaneous - 1×107 (1 dose) and intracranial 2x106 (1 dose) Mice treated with VAE showed significant median survival extension compared with mice treated with control r-HSV (∆ICP6, -/-γ34.5) or Endostar.65
RAMBO F ∆ICP6, -/-γ34.5, +Vstat120 U87ΔEGFR 1×105 (1 dose) The median survival of RAMBO was 28 days, which was statistically longer PBS-treated group (17 days).66
MGG23 1×105 (1 dose) The median survival of RAMBO was 65 days, which was significantly longer than mice treated with PBS.66
U87ΔEGFR 0.5×105 (1 dose) The median survival of RAMBO was 29 days, which was statistically longer than 19 days in PBS-treated group.147
GADD34 NG34 F ∆ICP6, -/-γ34.5, +GADD34 driven by nestin enhancer/promoter U87ΔEGFR-RliFluc and G35 and GL261N4 2x105 (1 dose) NG34 was shown to produce therapeutic efficacy as potent as rQnestin34.5 but with better tolerability.72
Anti-PD-1 NG34scFvPD-1 F ∆ICP6, -/-γ34.5, +GADD34 driven by nestin enhancer/promoter, +scFvPD-1 GL261N4 1.5x106 (1 dose) NG34scFvPD-1 treatment doubled the median survival time compared to non-anti-PD-1 expressing NG34 virus (69 vs 36.5 days).79
CT-2A 1.5x106 (1 dose) Neither NG34scFvPD-1 nor NG34 led to a significant increase in median survival compared to mock group. However, 2/12 mice treated with NG34scFvPD-1 survived for 120 days.79
CDH-1 OV-CDH1 F ∆ICP6, -/-γ34.5, +CDH-1 GBM30, Gl261N4 and U87ΔEGFR 2x105 (1 dose) OV-CDH1 significantly prolonged median survival compared to unarmed OV-Q1 in all three models.84
Chondroitinase ABC OV-Chase F ∆ICP6, -/-γ34.5, +Chase Gli36ΔEGFR–H2B-RFP and U87Δ EGFR Subcutaneous −106 v.p. (6–7 doses), intracranical - 3x105 v.p. (1 dose) In all models, treatment with OV-Chase significantly controlled tumor growth, resulted in extended survival compared to mock and unarmed oHSV.85
OV-ChaseM F ∆ICP6, -/-γ34.5, +ChaseM GBM30 3x105 v.p. (1 dose) The median survival of OV-ChaseM treated group was 33 days versus 22 days in group receiving PBS.88
PTENα HSV-P10 F ∆ICP6, -/-γ34.5, +PTENα, +eGFP U87Δ EGFR 1x105 (1 dose) HSV-P10 significantly prolonged median survival, resulted in and 30% long-term survivors compared to 10% long-term survivors in unarmed HSVQ treatment group.92
ULBP3 oHSVULBP3 F ∆ICP4, +miR-124, +ULBP3 XFM-Luc:PDGF,Cre 1x106 (1 dose) OHSVULBP3 inhibited tumor growth and prolonged median survival (18 days) compared to unarmed oHSV (8 days).95
oHSVULBP3-MMP9 F ∆ICP4, +miR-124, +ULBP3, +MMP9 XFM-Luc:PDGF,Cre 1x106 (1 dose) OHSVULBP3 showed statistically significant prolongation of median survival compared to OHSVULBP3-MMP9.96
ONCR-1 F ∆ICP4, -γ34.5, +miR-124, +ULBP3, +MMP9 U251 3x105-3x106 (1 dose) ONCR-1 inhibited tumor growth and prolonged survival compared to mock group.98
Pro-drug activating genes MGH2 F ∆ICP6, -/-γ34.5, -UL39, +CYP2B1, +shiCE Gli36ΔEGF 1.4×106 MGH2 did not prolong median survival compared to mock group.99

Abbreviations: PFU, plaque forming unit; (-), single deletion; (-/-), diploid deletion; ∆, mutation/inactivation; (+), insertion.