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Tertiary interactions formed by OLE*. (A) Base pairs observed at the 4H junction of OLE*. Nucleotides from stems P12.2, P13, P14, and P15 are colored in purple, green, blue, and red, respectively. The water molecule mediating the noncanonical C491-U573 base pair in stem P12.2 is shown as a red sphere. Hydrogen bonds are shown as gray dashed lines. (B–D) Minor groove base triple interactions formed between P12.2 helix and A504 (B), G503 (C), and U502 (D) in the P13 hairpin tetraloop.
