Table 2. Summary of the parameters probed by combined scanning SAXS and X-ray holography, including the nuclear structures accessible by each of the analyses and the physical property probed by the respective method.

Analysis approach	Nuclear structures accessed	Physical property probed
Dark field	Nucleoli, heterochromatin, euchromatin	Total scattering, Fourier components relevant for specific length scales
Power law exponent α	(Nucleoli)	Morphology
Porod constant K	Nucleoli, heterochromatin, euchromatin	Aggregation state, interface area
Phase shift	Nucleoli	Electron/mass density