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Details of the structural hierarchy of the major GQ structure formed in the human c-MYC promoter. (a) Guanine-rich sequences with four runs of at least three guanines self-associate into tetrads, consisting of a square planar arrangement of (b) Hoogsteen hydrogen-bonded guanines. (c) Monovalent metal ions, typically K+ (tan), stabilize the tetrad core and the highly ordered structure.
