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Longer (45–55 bp) linker lengths can better accommodate linker DNA stress in 1-start fibers. 0-start structures with straight DNA featured nucleosome breathing. The increased linker DNA significantly reduced the stress in linker DNA imposed by the 1-start stacking as compared to 167–177 NRL fibers. A large variety of nucleosome orientations was found in 2-start structures in which linker DNA was slightly deformed.
