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. 2021 Jan 25;134(4):1253–1262. doi: 10.1007/s00122-021-03771-9

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© The Author(s) 2021

Open AccessThis article is licensed under a Creative Commons Attribution 4.0 International License, which permits use, sharing, adaptation, distribution and reproduction in any medium or format, as long as you give appropriate credit to the original author(s) and the source, provide a link to the Creative Commons licence, and indicate if changes were made. The images or other third party material in this article are included in the article's Creative Commons licence, unless indicated otherwise in a credit line to the material. If material is not included in the article's Creative Commons licence and your intended use is not permitted by statutory regulation or exceeds the permitted use, you will need to obtain permission directly from the copyright holder. To view a copy of this licence, visit http://creativecommons.org/licenses/by/4.0/.

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Fig. 4 — Epistatic effects estimated in the lines with two or three QTLs for SER. a Plant type of HJX74 and pyramiding line P223. Scale bar, 15 cm. b Position of the three QTLs for SER in P223. Black part represents substitution segments from IR66897B (IB) with target QTLs, and white part is the genetic background of HJX74. c Epistatic effects estimated in S22, S33 and P223. S22 is a group of the SSSLs carrying qSER-2a and qSER-2b. S33 is a group of the SSSLs carrying qSER-3a and qSER-3b. P223 is a pyramiding line with qSER-2a, qSER-2b and qSER-3a. Student’s t test. SER stigma exsertion rate, SSSL single-segment substitution line