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. 2021 Jan 12;27(6):1250–1265. doi: 10.1111/gcb.15485

TABLE 1.

Predictor variables included in random‐forest analyses. All climate and weather variables were calculated from the 4 km PRISM daily data(PRISM Climate Group, 2019) for the year of bee sampling, 1 year prior to sampling, and 15‐year means (2001–2015; U.S. Geological Survey, 2014; USDA Douglas, Soba, et al., 2020, Douglas, Sponsler, et al., 2020; Koh et al., 2016; NASS, 2016)

Predictor variables Unit Source
Climate/Weather
Annual mean temperature °C BIOCLIM a
Mean diurnal range °C BIOCLIM a
Isothermality (temperature evenness) percent BIOCLIM a
Temperature seasonality (standard deviation × 100) °C BIOCLIM a
Maximum temperature of warmest month °C BIOCLIM a
Minimum temperature of coldest month °C BIOCLIM a
Temperature annual range °C BIOCLIM a
Mean temperature (wettest, driest, warmest, and coldest quarter) °C BIOCLIM a
Growing degree days (base 4.5 and 10°C) °C USDA NRCS a
Annual maximum and minimum temperature °C USDA NRCS a
Length of frost‐free growing season (−2.2 and 0°C threshold) days USDA NRCS a
Date of last spring frost (−2.2 and 0°C threshold) date USDA NRCS a
Date of first fall frost (−2.2 and 0°C threshold) date USDA NRCS a
Mean temperature in July °C USDA NRCS a
Monthly temperature range °C USDA NRCS a
Annual precipitation mm BIOCLIM a
Precipitation of wettest and driest month mm BIOCLIM a
Precipitation seasonality (coefficient of variation) mm BIOCLIM a
Precipitation (wettest, driest, warmest, and coldest quarter) mm BIOCLIM a
Growing season total precipitation mm USDA NRCS a
Mean number of days between 0.254 cm or greater rain events days USDA NRCS a
Mean number of days between 0.76 cm or greater rain events days USDA NRCS a
Monthly precipitation range cm USDA NRCS a
Topography
Elevation m USGS b
Slope percent USGS b
Winter and summer solar radiation watt hours/m 2 USGS b
Topographic convergence index NA USGS b
Aspect (degrees NS, EW) degrees USGS b
Landscape quality
Percent developed land within 1750 m radius percent USDA NASS c
Percent agriculture within 1750 m radius percent USDA NASS c
Percent natural land within 1750 m radius percent USDA NASS c
Floral resource quality (spring, summer, and fall) NA Koh et al d
Nesting resource quality (for spring, summer, and fall bees) NA Koh et al d
Combined resource quality (spring, summer, and fall) NA Koh et al d
Local habitat type NA USDA NASS c
Insecticide toxic load Douglas, Soba, et al. (2020), Douglas, Sponsler, et al. (2020)
Abundance of Apis mellifera in pan trap samples number of individuals BIML data
Sampling method
Volume and color of pan traps NA BIML data
a

PRISM Climate Group. http://prism.oregonstate.edu (2019).

b

U.S. Geological Survey. National Elevation Dataset. https://gdg.sc.egov.usda.gov/ (2014).

c

USDA NASS. USDA National Agricultural Statistics Service Cropland Data Layer. http://nassgeodata.gmu.edu/CropScape/ (2016).

d

Koh et al. Modeling the status, trends, and impacts of wild bee abundance in the United States. Proc. Natl. Acad. Sci. 113, 140–145 (2016).