Table 2:
Targeted Rickettsia screen of terrestrial invertebrates
| Terrestrial invertebrate group | Species | Location | Year | No. tested | No. positive |
|---|---|---|---|---|---|
| Araneae | Agelenopsis aperta | Tennessee, USA | N/A | 12 | 0 |
| Allopecosa pulverulenta | Berne, Germany | N/A | 16 | 0 | |
| Amaurobius fenestralis | Montpellier, France | 2006 | 16 | 1 | |
| Araneus diadematus | Beerse, Belgium | N/A | 19 | 0 | |
| Araneus diadematus | Greater London, UK | N/A | 8 | 0 | |
| Argiope bruennichi | Hamburg, Germany | N/A | 7 | 0 | |
| Argiope lobata | Spain | N/A | 7 | 0 | |
| Argiope lobata | Israel | N/A | 4 | 0 | |
| Cyclosa conica | Brandenburg, Germany | N/A | 11 | 0 | |
| Dysdera crocata | Montpellier, France | 2006 | 2 | 0 | |
| Enoplognatha ovata | Greater London, UK | N/A | 20 | 0 | |
| Erigone atra | Cheshire, UK | 2017 | 1 | 0 | |
| Evarcha falcata | Beerse, Belgium | N/A | 5 | 0 | |
| Holochnemus pluchei | Montpellier, France | 2006 | 7 | 0 | |
| Hylyphantes graminicola | Cheshire, UK | 2017 | 1 | 1 | |
| Larinioides cornutus | Greater London, UK | N/A | 6 | 0 | |
| Larinoides sclopetarius | Hamburg, Germany | N/A | 17 | 0 | |
| Linyphia triangularis | Berlin, Germany | N/A | 9 | 9 | |
| Linyphia triangularis | Greater London, UK | N/A | 6 | 0 | |
| Lycosa sp. | Cheshire, UK | 2017 | 2 | 0 | |
| Metellina mengei | Greater London, UK | N/A | 13 | 0 | |
| Metellina segmentata | Brandenburg, Germany | N/A | 9 | 0 | |
| Neriene clathrata | Beerse, Belgium | N/A | 13 | 0 | |
| Neriene peltata | Cheshire, UK | 2017 | 1 | 0 | |
| Pachygnatha degeeri | Berne, Germany | N/A | 11 | 0 | |
| Pachygnatha listeri | Beerse, Belgium | N/A | 17 | 0 | |
| Pardosa lugubris | Darmstadt, Germany | N/A | 20 | 1 | |
| Pardosa pullata | Brandenburg, Germany | N/A | 20 | 0 | |
| Pardosa purbeckensis | Belgium | N/A | 19 | 0 | |
| Pholcus phalangioides | Berlin, Germany | N/A | 20 | 17 | |
| Pisaura mirabilis | Greater London, UK | N/A | 12 | 1 | |
| Tetragnatha montana | Greater London, UK | N/A | 20 | 0 | |
| Tetragnatha sp. | Hampshire, UK | 2017 | 3 | 0 | |
| Unknown sp. | Cheshire, UK | 2017 | 2 | 0 | |
| Xysticus cristatus | Cambridgeshire, UK | N/A | 16 | 0 | |
| Opiliones | Leiobunum rotundum | Feurs, France | 2006 | 6 | 0 |
| Ixodida | Ixodes uriae | Hornøya, Norway | 2005 | 19 | 0 |
| Rhipicephalus microplus | New Caledonia | 2003 | 1 | 0 | |
| Scorpiones | Euscorpius flavicauda | St Nazaire de Pézan, France | 2006 | 1 | 0 |
| Diplopoda | Ommatoiulus sp. | Cheshire, UK | 2016 | 1 | 0 |
| Neuroptera | Unknown sp. | Cheshire, UK | 2017 | 1 | 0 |
| Mecoptera | Panorpa sp. | Cheshire, UK | 2017 | 2 | 0 |
| Orthoptera | Calliptamus italicus | Notre Dame de Londres, France | 2016 | 18 | 0 |
| Chorthippus brunneus | Uk | 2006 | 20 | 0 | |
| Gryllomorpha dalmatina | Montpellier, France | 2006 | 2 | 0 | |
| Blattaria | Loboptera decipiens | Montpellier, France | 2006 | 17 | 0 |
| Mantodae | Iris oratoria | St Nazaire de Pézan, France | 2006 | 6 | 0 |
| Mantis religiosa | Feurs, France | 2006 | 3 | 0 | |
| Dermaptera | Forficula Auricularia | Feurs, France | 2006 | 9 | 0 |
| Hemiptera | Aphis fabae | Montpellier, France | 2006 | 12 | 0 |
| Aphis nerii | Montpellier, France | 2006 | 8 | 0 | |
| Baizongia pistaciae | Viols le Fort, France | 2006 | 12 | 0 | |
| Cicadella viridis | L'Olme, France | 2006 | 16 | 0 | |
| Cimex lectularius | Yorkshire, UK | 2008 | 12 | 12 | |
| Elasmucha grisea | Greater London, UK | 2006 | 16 | 0 | |
| Graphosoma italicum | Montpellier, France | 2006 | 12 | 0 | |
| Lygaeus equestris | Montpellier, France | 2006 | 12 | 0 | |
| Notostira elongata | L'Olme, France | 2006 | 11 | 0 | |
| Pyrrhocoris apterus | Montpellier, France | 2006 | 11 | 0 | |
| Rhyparochromus vulgaris | Castelnaudary, France | 2006 | 20 | 0 | |
| Coleoptera | Anaspis frontalis | Mont Barri, France | 2004 | 12 | 0 |
| Anthaxia nitidula | Mont Barri, France | 2004 | 20 | 0 | |
| Anthaxia sp. | Mont Barri, France | 2004 | 16 | 0 | |
| Calvia 14-guttata | Greater London, UK | 2006 | 6 | 0 | |
| Capnodis tenebrionis | Montpellier, France | 2006 | 1 | 0 | |
| Cetonia aurata | Feurs, France | 2006 | 3 | 0 | |
| Cetonia aurata | Mont Barri, France | 2004 | 12 | 0 | |
| Chrysolina varians | Mont Barri, France | 2004 | 18 | 0 | |
| Clytus arietis | Mont Barri, France | 2004 | 20 | 0 | |
| Dermestes sp. | Mont Barri, France | 2004 | 20 | 0 | |
| Dermestes tessellatocollis | Cheshire, UK | 2016 | 2 | 0 | |
| Gastrophysa sp. | Greater London, UK | 2006 | 20 | 0 | |
| Geotrupes stercorarius | Mont Barri, France | 2004 | 3 | 0 | |
| Larinus scolymi | Aldira de Irmeros, Spain | 2005 | 12 | 0 | |
| Leptinotarsa decemlineata | Feurs, France | 2006 | 10 | 0 | |
| Mordellistena sp. | Mont Barri, France | 2004 | 10 | 0 | |
| Oedemera sp. | Mont Barri, France | 2004 | 20 | 0 | |
| Oncocerna sp. | Mont Barri, France | 2004 | 20 | 0 | |
| Phyllobius argentatus | Mont Barri, France | 2004 | 15 | 4† | |
| Pseudovadonia livida | Mont Barri, France | 2004 | 19 | 0 | |
| Stenopterus sp. | Mont Barri, France | 2004 | 20 | 0 | |
| Diptera | Braula coeca | Ouessant, France | 2002 | 4 | 0 |
| Chorisops tunisiae | Montpellier, France | 2003 | 8 | 0 | |
| Delia antiqua | N/A | N/A | 11 | 0 | |
| Delia platura | N/A | N/A | 11 | 0 | |
| Delia radiacum | N/A | N/A | 10 | 0 | |
| Gasterophilus intestinalis | France | N/A | 10 | 0 | |
| Hippobosca equina | Restinclières, France | 2006 | 15 | 0 | |
| Lonchoptera lutea | Cheshire, UK | 2017 | 3 | 0 | |
| Medetera petrophila | St Bauzille de Putois, France | 2003 | 12 | 0 | |
| Musca domestica | L'Olme, France | 2006 | 20 | 0 | |
| Musca vitripennis | Notre Dame de Londres, France | 2003 | 8 | 0 | |
| Neomyia cornicina | Notre Dame de Londres, France | 2003 | 8 | 0 | |
| Protocalliphora sp. | Corse, France | 2003 | 2 | 0 | |
| Protocalliphora azurea | Montpellier, France | 2005 | 12 | 12 | |
| Psila rosae | N/A | N/A | 11 | 0 | |
| Stomoxys calcitrans | Le Malzieu, France | 2001 | 11 | 0 | |
| Lepidoptera | Chilo phragmitellus | Feurs, France | 2006 | 10 | 0 |
| Euplagia quadripunctaria | Feurs, France | 2006 | 2 | 0 | |
| Pieris brassicae | Feurs, France | 2006 | 7 | 0 | |
| Plodia interpunctella | Montpellier, France | 2006 | 12 | 0 | |
| Thymelicus lineola | Greater London, UK | 2006 | 15 | 0 | |
| Thymelicus sylvestris | Greater London, UK | 2006 | 2 | 0 | |
| Triodia sylvina | Montpellier, France | 2006 | 4 | 0 | |
| Hymenoptera | Amblyteles armatorius | St Nazaire de Pézan, France | 2006 | 1 | 0 |
| Amegilla albigena | St Nazaire de Pézan, France | 2006 | 13 | 0 | |
| Amegilla ochroleuca | St Nazaire de Pézan, France | 2006 | 3 | 0 | |
| Anthidium florentinum | St Nazaire de Pézan, France | 2006 | 6 | 0 | |
| Apis mellifera | UK | 2006 | 9 | 0 | |
| Bombus terrestris | Northwest Switzerland | 2006 | 20 | 0 | |
| Diplolepis rosae | L'Olme, France | 2006 | 2 | 0 | |
| Formica lugubris | UK | 2006 | 10 | 0 | |
| Pachycrepoideus sp. | UK | N/A | 94 | 6‡ | |
| Polistes dominulus | St Nazaire de Pézan, France | 2006 | 4 | 0 | |
| Polistes nimpha | St Nazaire de Pézan, France | 2006 | 19 | 0 | |
| Sceliphron caementarium | St Nazaire de Pézan, France | 2006 | 3 | 0 |
A species was deemed positive through PCR and designated to Rickettsia group after Sanger sequencing and phylogenetic placement. All strains belong to the Torix group except † = Rhyzobius and ‡ = Belli. Bold entries indicate Rickettsia-positive hosts identified in this study. N/A: not available.