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. 2021 Feb 26;11(3):354. doi: 10.3390/biom11030354

Table 2.

PERK pathway modulators.

Modulators (Compounds) PERK Pathway Outcome Neurodegenerative Diseases Additional Targets
MK-28 Activation
(via activating PERK)
HD [87]
CCT020312 Activation
(Nrf2 branch)
PSP [129]
GSK260414 Inhibition
(via inhibiting PERK)
Prion Disease [130], PD [131], FD [132], AD [133] RIPK1 [134]
GSK2656157 Traumatic brain
injury [135]
Echinacoside (ECH) AD [136,137], PD [138] Ghrelin receptor [139], Androgen
receptor [140]
SC79 Inhibition
(activates AKT causing inhibitory phosphorylation of PERK kinase loop)
Prion Disease [141]
2BAct Inhibition
(downstream of P-eIF2α, via eIF2B activation)
VWMD [58]
ISRIB ALS [142], VWMD [59], AD [143], TBI [144]
Dibenzoylmethane
(DBM)
Inhibition
(downstream of P-eIF2α, similar activity to that of ISRIB)
FTD [145], Prion disease [145], PD [146] Nrf2 [147], AMPK [148]
Trazodone HCl T-type calcium channel [149], monoamine receptor [150]
Guanabenz Activation
(via inhibiting dephosphorylation of P-eIF2α, inhibits GADD34)
VWMD [117], ALS [116], PD [118,120] ASICs [151]
PromISR-6
(guanabenz analog, target unknown)
HD [127] Possibly activator of PERK and other eIF2α kinases [127]
Salubrinal PD [53,56,114], TBI [113], HD [79] CReP inhibitor, Dusp2 (PAC1) inhibitor [152]
Sephin1 ALS [123], Charcot-Marie-Tooth disease [123], MS [124], Prion Disease [125] ASICs [151]
Raphin1 Activation
(via inhibiting dephosphorylation of P-eIF2α, inhibits CReP)
HD [128]
Bruceine D Activation
(Nrf2 branch, mechanism unknown)
PD [153] Notch [154],
JNK [155]
Naringenin (NAR) PD [156], AD [157,158] CRMP-2 [158]
Sulforaphane (SFN) Activation
(Nrf2 branch—acts on KEAP1, releasing Nrf2)
PD [159,160,161], HD [162], AD [163,164,165,166], MS [167], FRDA [168] BACE1 [169],
NF-κB [170]
Tertbuthyl-hydroquinene (tBHQ) Activation
(Nrf2 branch)
HD [171], AD [172], PD [173]
Acetylenic tricyclic bis(cyanoenone) TBE-31 Activation
(Nrf2 branch—binds KEAP1, thus releasing Nrf2)
FRDA [168]
RTA-408
(omaveloxolone)
FRDA [174]
Dimethylfumarate (DFM) MS [175], PD [176,177] MSK1 and RSK1 [178,179] and others [180]