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T4SS of Brucella is a multiprotein complex encoded by VirB operon, which participates in the intracellular activities of Brucella. The T4SS is mainly divided into the following five parts: the elongation region is composed of VirB2; the central and outer membrane regions were composed of VirB7, VirB9 and VirB10; the junction region is composed of VirB5 and VirB10; the intimal region was composed of VirB3, VirB4, VirB6, VirB8 and VirB10; ATP energy region composed of VirB4 and VirB11.
