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. 2021 Apr 16;11:8338. doi: 10.1038/s41598-021-87495-2

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© The Author(s) 2021

Open Access This article is licensed under a Creative Commons Attribution 4.0 International License, which permits use, sharing, adaptation, distribution and reproduction in any medium or format, as long as you give appropriate credit to the original author(s) and the source, provide a link to the Creative Commons licence, and indicate if changes were made. The images or other third party material in this article are included in the article's Creative Commons licence, unless indicated otherwise in a credit line to the material. If material is not included in the article's Creative Commons licence and your intended use is not permitted by statutory regulation or exceeds the permitted use, you will need to obtain permission directly from the copyright holder. To view a copy of this licence, visit http://creativecommons.org/licenses/by/4.0/.

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Gular region of selected hyperoliid taxa. (a) Congolius robustus gen. et comb. n. (IVB-H-CD18-255) with inflated vocal sac, (b) Hyperolius balfouri (IVB-H-CD15-062) with inflated vocal sac, (c) C. robustus gen. et comb. n. (ZMUC-R.771175), (d) Morerella cyanophthalma (ZMB 71588, paratype), (e) Cryptothylax greshoffii (NMP-P6V 76076/1), (f) H. balfouri (IVB-H-CD15-061), (g) H. cf. cinnamomeoventris (IVB-H-CD18-092), (h) H. phantasticus (IVB-H-CD18-203), (i) H. cf. adspersus (IVB-H-CD15-139), (j) Chrysobatrachus cupreonitens (MCZ A-64739, paratype), (k) Callixalus pictus (MCZ A-64707), (l) Afrixalus equatorialis (NMP-P6V 76073/1), (m) Opisthothylax immaculatus (NMP-P6V 76090/1), (n) Phlyctimantis verrucosus (NMP-P6V 76085/1). Photos by (a,b,j,k) V.G., (c,e–i,l–n) T.N. and (d) F. Tillack and M.-O. Rödel.