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. 2021 May 6;11:9661. doi: 10.1038/s41598-021-88834-z

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© The Author(s) 2021

Open Access This article is licensed under a Creative Commons Attribution 4.0 International License, which permits use, sharing, adaptation, distribution and reproduction in any medium or format, as long as you give appropriate credit to the original author(s) and the source, provide a link to the Creative Commons licence, and indicate if changes were made. The images or other third party material in this article are included in the article's Creative Commons licence, unless indicated otherwise in a credit line to the material. If material is not included in the article's Creative Commons licence and your intended use is not permitted by statutory regulation or exceeds the permitted use, you will need to obtain permission directly from the copyright holder. To view a copy of this licence, visit http://creativecommons.org/licenses/by/4.0/.

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A model of SEs maturation regulatory network in two different media. Differences in shoot and root apical meristems observed in the SEs are related to expression changes of key genes involved in cytokinin signaling, stem cell maintenance and embryogenesis. Initial samples were SEs from PEM at globular and early torpedo steges. The second sample were secondary SEs at cotyedonary stage in the second month of maturation. SEs were cleary developed with specific shape in osmotic stress medium contrasting with non-osmotis stress medium.