Fig. 3.

(A) Streak image of cell motion. Raw images are obtained at cell density $178\hspace{0.17em}{\mathrm{m}\mathrm{m}}^{-\mathrm{2}}$ and averaged over 10 s to produce the streak image. (Scale bar, $200\hspace{0.17em}\mathrm{\mu }\text{m}$.) (B) Instantaneous cell coordinates (gray line, $\mathit{r}\left(t\right)$) are averaged over a sliding window of $2\hspace{0.17em}\mathrm{s}$ to highlight the noncircular motion (green line, $\overline{\mathit{r}}\left(t\right)$). (C) Temporal history of instantaneous (gray) and window-averaged (green) coordinates. (D) Probability distribution functions (PDF) of window-averaged cell displacements (squared and normalized by time separation, $s\equiv △{\overline{r}}^2/4△t$) from experiment (circles) and model (squares) for four time separations. The experiment and model results are in agreement; for presentation clarity, we rescale model results in D by a factor of 100.