MODEL 2 (fast MSO neurons, 600-Hz channels): lateralisation of anechoic and reverberant speech ‘church’, with and without adaptation of synaptic inputs from auditory nerve to CN: (a) waveforms and ITDs of anechoic speech, gammatone filtered at 600 Hz (top row); mean firing rates in left MSO (middle row, red) and right MSO (middle row, blue); AUCtrial neurometric (bottom row). Anechoic speech is correctly lateralised to the right (pink vertical bands) with nonadapting (left column) or adapting synapses (right column). (b) Reverberant speech produces highly variable ITDs and confounding ILDs (top row), and requires adapting synapses (right column) for consistently correct lateralisations (pink vertical bands). Nonadapting synapses (left column) produced both correct (pink vertical bands) and incorrect lateralisations (light blue vertical bands). Synaptic adaptation removed all the incorrect lateralisations, leaving correct lateralisations at stimulus onsets