Figure 4.

Transition rate matrix for the dependent model of evolution between brush organs and scent pads, suggesting that gains are unlikely to occur when males of a species already possess one or the other type of androconial organ. Density plots depict the transition rates for discrete correlated traits inferred using Pagel's method [40] as implemented in Phytools [39] fitted to 1000 randomly sampled trees from the bootstrap distribution used to estimate the support for the consensus tree. Arrow width (approximately) scaled to average rate estimates. Dashed arrows indicate transition rates close to zero based on the ML and Bayesian analyses implementing uncertainty. (Online version in colour.)