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. 2021 Apr 30;12:626715. doi: 10.3389/fmicb.2021.626715

TABLE 2.

Gene expression response of PA14 grown in the presence of tobramycin versus untreated control.

Gene name Gene ID Gene product RNA-seq Ribo-seq
FC1 FC
Energy metabolism and tricarboxylic acid cycle cycle
PA14_06800 PA14_06800 Hypothetical protein 18.422 X3
ldh PA14_19870 Leucine dehydrogenase –13.38 –2.23
PA14_19900 PA14_19900 Pyruvate dehydrogenase E1 component subunit alpha –104.77 –4.45
pdhB PA14_19910 Pyruvate dehydrogenase E1 component. beta chain –94.26 –2.53
PA14_19920 PA14_19920 Branched-chain alpha-keto acid dehydrogenase subunit E2 –78.9 X
nqrA PA14_25280 Na(+)-translocating NADH-quinone reductase subunit A 6.06 ND4
nqrB PA14_25305 Na(+)-translocating NADH-quinone reductase subunit B ND –4.6
nqrC PA14_25320 Na(+)-translocating NADH-quinone reductase subunit C –2.08 –2.66
nqrD PA14_25330 Na(+)-translocating NADH-quinone reductase subunit D –5.14 –3.25
nqrE PA14_25340 Na(+)-translocating NADH-quinone reductase subunit E –7.06 –2.51
nqrF PA14_25350 Na(+)-translocating NADH-quinone reductase subunit F –9.6 ND
nuoN PA14_29850 NADH dehydrogenase subunit N –20.61 –13.71
nuoM PA14_29860 NADH dehydrogenase subunit M –24.8 –3.99
nuoL PA14_29880 NADH dehydrogenase subunit L –25.56 –3.4
nuoK PA14_29890 NADH dehydrogenase subunit K –8 –6.97
nuoJ PA14_29900 NADH dehydrogenase subunit J –5.16 –15.98
nuoI PA14_29920 NADH dehydrogenase subunit I –9.55 –13.6
nuoH PA14_29930 NADH dehydrogenase subunit H –10 –4.88
nuoG PA14_29940 NADH dehydrogenase subunit G –46.39 –8.11
nuoF PA14_29970 NADH dehydrogenase I subunit F –13.36 –8.16
nuoE PA14_29980 NADH dehydrogenase subunit E –13.55 –5.35
icd PA14_30190 Iisocitrate dehydrogenase –3.09 ND
sucD PA14_43940 Succinyl-CoA synthetase subunit alpha –9.39 –3.27
sucC PA14_43950 Succinyl-CoA synthetase subunit beta –3.73 ND
lpdG PA14_43970 Dihydrolipoamide dehydrogenase –16.47 –5.26
sucB PA14_44000 Dihydrolipoamide succinyltransferase –10.98 –9.15
sucA PA14_44010 2-oxoglutarate dehydrogenase E1 –4.27 –4.12
PA14_53970 PA14_53970 Aconitate hydratase –18.05 –8.09
Phenylalanine/Tyrosine catabolism
fahA PA14_38530 Fumarylacetoacetase –30.98 –2.32
maiA PA14_38550 Maleylacetoacetate isomerase –48.44 –3.57
phhB PA14_53000 Pterin-4-alpha-carbinolamine dehydratase –14.76 –2.47
phhC PA14_53010 Aromatic amino acid aminotransferase –13.94 –2.78
Arginine catabolism
arcD PA14_68300 Arginine/ornithine antiporter –49.29 2.29
arcA PA14_68330 Arginine deiminase –77.18 –2.12
arcB PA14_68340 Ornithine carbamoyltransferase –137.35 –4.19
Leucin/Valine/Isoleucin degradation and biosynthesis
lpdV PA14_35490 Dihydrolipoamide dehydrogenase –104.5 –6.56
bkdB PA14_35500 Branched-chain alpha-keto acid dehydrogenase subunit E2 –91.67 –5.51
bkdA2 PA14_35520 2-oxoisovalerate dehydrogenase subunit beta –104.31 –4.84
bkdA1 PA14_35530 2-oxoisovalerate dehydrogenase subunit alpha –5.23 X
gnyR PA14_38430 Regulatory gene of gnyRDBHAL cluster. GnyR –12.88 ND
gnyD PA14_38440 Citronelloyl-CoA dehydrogenase. GnyD –30.42 –2.32
gnyB PA14_38460 Acyl-CoA carboxyltransferase subunit beta –27.71 X
gnyH PA14_38470 Gamma-carboxygeranoyl-CoA hydratase –33.85 –3.73
gnyA PA14_38480 Alpha subunit of geranoyl-CoA carboxylase. GnyA –26.11 X
ilvA2 PA14_47100 Threonine dehydratase 24.7 7.68
Peptidoglycan biosynthesis
ddl PA14_57320 D-alanine–D-alanine ligase –151.66 –8.63
murC PA14_57330 UDP-N-acetylmuramate–L-alanine ligase –78.75 –9.99
murG PA14_57340 UDPdiphospho-muramoylpentapeptide beta-N-acetylglucosaminyl transferase –44.2 –8.27
murD PA14_57370 UDP-N-acetylmuramoyl-L-alanyl-D-glutamate synthetase –46.8 –10.56
mraY PA14_57380 Phospho-N-acetylmuramoyl-pentapeptide-transferase –13.67 –2.18
murF PA14_57390 UDP-N-acetylmuramoylalanyl-D-glutamyl-2.6-diaminopimelate–D-alanyl-D-alanine ligase –11.12 X
murE PA14_57410 UDP-N-acetylmuramoylalanyl-D-glutamate–2. 6-diaminopimelate ligase –55.53 –9.29
rmlC PA14_68210 dTDP-4-dehydrorhamnose 3.5-epimerase –12.92 –3.34
Glycogen metabolism
glgA PA14_36570 Glycogen synthase –3.09 –5.06
PA14_36580 PA14_36580 Glycosyl hydrolase –19.03 –11.25
PA14_36590 PA14_36590 4-alpha-glucanotransferase –34.5 –21.54
PA14_36605 PA14_36605 Maltooligosyl trehalose synthase –27.97 –10.78
PA14_36620 PA14_36620 Hypothetical protein –44.57 –10.58
PA14_36630 PA14_36630 Glycosyl hydrolase –25.56 –6.53
glgB PA14_36710 Glycogen branching protein –38.34 –15.21
PA14_36730 PA14_36730 Trehalose synthase –30.78 –16.68
PA14_36740 PA14_36740 Hypothetical protein –4.77 –5.04
glgP PA14_36840 Glycogen phosphorylase –5.98 –2.93
PA14_36850 PA14_36850 Hypothetical protein –2.26 –2.9
Pathogenicity and virulence
tssL1 PA14_00925 Hypothetical protein –17.71 X
tssk1 PA14_00940 Hypothetical protein –6.38 –4.32
tssJ1 PA14_00960 Lipoprotein –12.26 –3.67
PA14_00960 PA14_00970 Hypothetical protein –33.07 –2.84
pilJ PA14_05360 Twitching motility protein PilJ –2.63 ND
pilK PA14_05380 Methyltransferase PilK –2.59 ND
chpA PA14_05390 ChpA –11.54 –4.98
PA14_05400 PA14_05400 Methylesterase –20.38 –3.72
PA14_34000 PA14_34000 HsiH3 –23.95 –5.54
stk1 PA14_42880 Stk1 –2.19 X
stp1 PA14_42890 Stp1 –5.56 –3.6
PA14_42900 PA14_42900 IcmF2 –3.59 –2.57
PA14_42910 PA14_42910 DotU2 –11.99 –3.08
PA14_42920 PA14_42920 HsiJ2 –7.75 –3.42
PA14_42940 PA14_42940 Lip2.2 –20.14 –3.56
PA14_42950 PA14_42950 Fha2 –25.63 –5.86
PA14_42960 PA14_42960 Lip2.2 –77.39 X
PA14_42970 PA14_42970 Sfa2 –13.89 –6.03
PA14_42980 PA14_42980 ClpV2 –10.86 –5.68
PA14_42990 PA14_42990 HsiH2 –10.56 –4.18
PA14_43000 PA14_43000 HsiG2 –14.05 –3.6
PA14_43020 PA14_43020 Hypothetical protein –10.04 –3.57
PA14_43030 PA14_43030 HsiC2 –4.85 X
flhB PA14_45720 Flagellar biosynthesis protein FlhB –5.62 X
fliR PA14_45740 Flagellar biosynthesis protein FliR –8.13 X
fliQ PA14_45760 Flagellar biosynthesis protein FliQ –10.6 X
fliP PA14_45770 Flagellar biosynthesis protein FliP –6.41 X
fliN PA14_45790 Flagellar motor switch protein –2.05 X
flgK PA14_50360 Flagellar hook-associated protein FlgK –75.19 –3.47
flgJ PA14_50380 Flagellar rod assembly protein/muramidase FlgJ –9.17 –4.58
flgI PA14_50410 Flagellar basal body P-ring protein –6.22 X
flgH PA14_50420 Flagellar basal body L-ring protein –3.99 X
flgG PA14_50430 Flagellar basal body rod protein FlgG ND 2.13
flgF PA14_50440 Flagellar basal body rod protein FlgF ND 3.54
pqsE PA14_51380 Quinolone signal response protein –80.7 –2.98
pqsD PA14_51390 3-oxoacyl-ACP synthase –90.01 –5.86
pqsC PA14_51410 PqsC –44.86 –4.06
pqsB PA14_51420 PqsB –20.71 –2.34
pqsA PA14_51430 PqsA –3.73 X
PA14_55780 PA14_55780 Phosphate transporter –46.70 X
PA14_55790 PA14_55790 Two-component sensor –15.49 –2.70
PA14_55800 PA14_55800 Hypothetical protein –2.21 X
PA14_55810 PA14_55810 Hypothetical protein –2.73 –2.00
PA14_55820 PA14_55820 Two-component response regulator –25.16 X
PA14_55840 PA14_55840 Hypothetical protein –84.76 X
PA14_55850 PA14_55850 Hypothetical protein –68.52 X
PA14_55860 PA14_55860 Pilus assembly protein –98.84 X
PA14_55880 PA14_55880 Hypothetical protein –105.85 X
cpaF2 PA14_55890 Hypothetical protein –111.85 X
PA14_55900 PA14_55900 Type II secretion system protein –36.82 –9.16
PA14_55920 PA14_55920 Hypothetical protein –15.84 –2.00
PA14_55930 PA14_55930 Type II secretion system protein –2.75 ND
PA14_55940 PA14_55940 Pilus assembly protein –8.25 –7.96
pilC PA14_58760 Type 4 fimbrial biogenesis protein pilC –2.41 ND
pilD PA14_58770 Type 4 prepilin peptidase PilD –11.48 ND
coaE PA14_58780 Dephospho-CoA kinase ND 2.48
fimU PA14_60280 Type 4 fimbrial biogenesis protein FimU –2.46 ND
pilW PA14_60290 Type 4 fimbrial biogenesis protein PilW –11.99 X
pilX PA14_60300 Type 4 fimbrial biogenesis protein PilX –9.12 X
pilY1 PA14_60310 Type 4 fimbrial biogenesis protein PilY1 –4.06 ND
pilE PA14_60320 Type 4 fimbrial biogenesis protein PilE –3.43 –2.53
PA14_65520 PA14_65520 Hypothetical protein –21.34 –3.44
PA14_65540 PA14_65540 Hypothetical protein –4.26 X
estA PA14_67510 Esterase EstA –11.29 –2.17
ABC transporters and Sha antiporter
opuCA PA14_13580 ABC transporter ATP-binding protein –28.83 –7.05
opuCB PA14_13590 ABC transporter permease –6 –4.83
opuCD PA14_13600 ABC transporter substrate-binding protein –4.73 –4.41
nppA2 PA14_41130 ABC transporter substrate-binding protein NppA2 –1.97 X
nppB PA14_41140 Peptidyl nucleoside antibiotic ABC transporter permease NppB –10.23 X
nppC PA14_41150 Peptidyl nucleoside antibiotic ABC transporter permease NppC –30.15 –3.71
nppD PA14_41160 Peptidyl nucleoside antibiotic ABC transporter ATP-binding protein NppD –22.25 –4.98
fabI PA14_41170 NADH-dependent enoyl-ACP reductase –21.89 –5.27
phaG PA14_50680 ShaA –44.11 –3.99
phaF PA14_50690 ShaB –29.33 –3.77
phaE PA14_50700 ShaC –12.23 –2.9
phaD PA14_50710 ShaD –7.08 –5.88
phaC PA14_50720 ShaE –8.48 –7.78
dppC PA14_58450 Dipeptide ABC transporter permease DppC –13.03 –3.55
dppD PA14_58470 Dipeptide ABC transporter ATP-binding protein DppD –34.43 –10.45
dppF PA14_58490 Dipeptide ABC transporter ATP-binding protein DppF –21.5 –4.19
Transcription and translation
tufB PA14_08680 Elongation factor Tu 92.01 3.09
rplC PA14_08850 50S ribosomal protein L3 27.97 ND
rplD PA14_08860 50S ribosomal protein L4 23.35 2.05
tyrS PA14_10420 Tyrosyl-tRNA synthetase 37.72 11.48
orf2 PA14_12350 (dimethylallyl)adenosine tRNA methylthiotransferase 23.15 2.17
rimM PA14_15980 16S rRNA-processing protein RimM 55.59 5.82
trmD PA14_15990 tRNA (guanine-N(1)-)-methyltransferase 52.24 4.74
rpsB PA14_17060 30S ribosomal protein S2 106.77 2.27
deaD PA14_27370 ATP-dependent RNA helicase 122.76 2.02
infC PA14_28660 Translation initiation factor IF-3 12.93 2.59
yadB PA14_62510 Glutamyl-Q tRNA(Asp) synthetase 34.45 4.5
yhbC PA14_62780 Hypothetical protein 14.37 ND
smpB PA14_63060 SsrA-binding protein 12.7 2.36
rpmE PA14_66710 50S ribosomal protein L31 316.32 ND
prfH PA14_72200 Peptide chain release factor-like protein 491.76 5.65
rnpA PA14_73420 Ribonuclease P 164.46 16.33
Stringent response and toxin-antitoxin systems
PA14_01510 PA14_01510 Hypothetical protein 25.78 4.87
PA14_01520 PA14_01520 Hypothetical protein 28.84 4.71
ndk PA14_14820 Nucleoside diphosphate kinase 6.47 ND
obgE PA14_60445 GTPase ObgE 15.46 2.42
vapI PA14_61840 Antitoxin HigA 9.67 5.8
rnk PA14_69630 Nucleoside diphosphate kinase regulator 11.23 2.08
spoT PA14_70470 Guanosine-3′.5′-bis(diphosphate) 3′-pyrophosphohydrolase 9.2 3.64

1FC, fold-change, p-value ≤ 0.05. 2Genes with FC ≤ –10 or ≥10 are represented in bold. 3X, not efficiently translated in the control and antibiotic treated samples, Ribo-seq BaseMean ≤ 50. 4ND, not differentially expressed, –2 ≤ FC ≤ 2 and/or p-value ≥ 0.05.