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. 2021 Mar 31;121(6):1543–1566. doi: 10.1007/s00421-021-04664-x

Table 4.

Resting haematological characteristics of breath-hold divers

References Participants Performance characteristics Red blood cells Haemoglobin Haematocrit Reticulocytes
Kang et al. (1963) 20 female habitual BHD (Ama) and 20 female ND

BHD:3.53 ± 0.05 × 106/mm3

ND: 3.63 ± 0.12 × 106/mm3

BHD: 12.8 ± 0.2 g/100 ml*

ND: 11.9 ± 0.3 g/100 ml

BHD: 35.1 ± 0.6%

ND: 34.6 ± 0.7%

Ferretti et al. (1991) 1 male (age: 57 years) and 2 female EBHD (mean age: 29 years)

PB Depth No limits category:

Male 101 m, and female 70 m and 80 m

Male: 148 g/L

Female: 143 g/L and

137 g/L

Qvist et al. (1993) 5 female Korean Ama (age: 38–52 years) Perform repeated dives to depths of 3–7 m for up to 4 h/day, spending 25–30% of this time underwater and completing > 100 dives during a single workday 12.8 ± 0.3 g/dl 38.5 ± 0.6%
Baković et al. (2003) 10 male trained BHD (age: 28.6 ± 1.7 years) and 10 male untrained individuals (age: 27.8 ± 2.4 years) Four were national team members

BHD: 41.3 ± 0.01%

Untrained: 43.8 ± 0.01%

Richardson et al. (2005) 29 male and 4 female EBHD (age: 30.3 ± 10.6 years), 23 elite male skiers (age: 20.2 ± 3 years), 23 male and 9 female untrained individuals (age: 30 ± 6.3 years)

Training background:

5.5 ± 4 h per week

EBHD: 150.1 g/L

Skiers: 145.5 g/L

Untrained: 146.9 g/L

Bakovic et al. (2005) 18 trained male BHD and 18 male ND Years practicing apnoea: 9.5 years (4–26 years)

BHD: 147.7

(125–177 g/L)

ND: 147.9

(131–172 g/L)

BHD: 43% (36–53%)

ND: 43% (39–49%)

Stewart et al. (2005) 10 trained BHD (age: 36 ± 11.3 years) and 10 ND (age: 32 ± 9.5 years) Years practicing apnoea: 17 years

BHD: 14.4 ± 1.6 g/dl

ND: 14.2 ± 0.9 g/dl

BHD: 44.3 ± 5.6%

ND: 40.6 ± 4.4%

Prommer et al. (2007) 7 male (age: 35 ± 9 years) and 3 female trained BHD (age: 32 ± 6 years), 7 male Scuba divers (age: 38 ± 11 years), 10 male untrained ND (age: 31 ± 10 years) and 9 non-diving triathletes (age: 31 ± 8 years)

Training background:

2–3 h, 3 times per week or more over at least the 3 years

BHD male: 152 ± 5 g/L

BHD female: 141 ± 9 g/L

Scuba divers: 154 ± 9 g/L

Untrained ND: 151 ± 8 g/L

Triathletes: 152 ± 10 g/L

Schagatay et al. (2007) 7 male BHD (age mean: 27 years [22–35 years]) 14.18 ± 0.48 g/dl 43.5 ± 1.3%
Kjeld et al. (2009) 8 male and 1 female trained competitive BHD, 5 male and 1 ND Current or former national record holders 142 g/L
Richardson et al. (2009) 10 male BHD

Training background:

 < 2 h per week

134.1 ± 0.8 g/L 40.9 ± 2%
Sureda et al. (2015) 7 male professional spearfishing BHD (age: 30 ± 2.1 years)

Years practicing apnoea:

5—20 years

Training background:

3–4 h diving 5 times per week

4.93 ± 0.07 × 1012/L 14.7 ± 0.1 g/dl 42.8 ± 0.4%
Oh et al. (2017) 715 female Haenyeo habitual BHD and 715 matched ND

BHD: 13.0 ± 1.2 g/dl

ND: 13.0 ± 1.1 g/dl

Fernandez et al. (2017) 56 male BHD (age: 36.6 ± 8.5 years [22–57 years])

Years practicing apnoea:

2.09 ± 0.7 years

15.6 ± 1.1 g/dl

(14.1—17.7 g/dl)

Kjeld et al. (2018) 8 male BHD (age: 42 ± 8 years) and 8 male judo athletes (age: 36 ± 11years)

PB STA:

403 ± 62 s

PB DYN:

163 ± 36 m

PB DNF:

133 ± 37 m

BHD: 15.2 ± 0.8 g/dl

ND: 15.3 ± 0.8 g/dl

Puspitaningrum et al. (2019) 21 male habitual BHD (Bajau) (age: 15–62 years)

15.62 g/dl

(12.9–16.9 g/dl)

Elia et al. (2019b) 11 male EBHD and 10 male matched control ND

Years practicing apnoea:

7 ± 2 years

Training background:

8 ± 2 h per week

PB STA:

414 ± 101 s

PB DYN:

202 ± 46 m

PB DNF:

145 ± 50 m

EBHD: 5.43 ± 0.49 •1012/L*

(4.80–6.3 × 1012/L)

ND: 4.93 ± 0.24 × 1012/L

(4.64–5.34 × 1012/L)

EBHD: 153 ± 6 g/L*

(144–161 g/L)

ND: 148 ± 3 g/L

(145–152 g/L)

EBHD: 46 ± 4%

(42–52%)

ND: 45 ± 2%

(43–49%)

EBHD: 1.47 ± 0.29%*

(1.1–2%)

ND: 1.13 ± 0.30%

(0.7–1.6%)

Marlinge et al. (2019) 12 male experienced BHD (age: 37 ± 8 years), 12 male experienced Scuba divers (age: 51 ± 10 years) and 8 male ND (age: 48 ± 4 years)

BHD: 15 ± 0.3 g/dl

Scuba: 15.2 ± 1.5 g/dl

Control: 15.3 ± 0.9 g/dl

BHD: 45 ± 0.9%

Scuba: 44.9 ± 1.2%

Control: 45.5 ± 1.8%

Fernandez et al. (2019) 29 male BHD (age: 36 ± 5 years) Years practicing apnoea: mean of 2 years

15.9 ± 1.2 g/dl

15.4 ± 1.2 g/dl

15.5 ± 0.8 g/dl

Elia et al. (2021b) 8 male EBHD and 10 male ND

Years practicing apnoea:

6 ± 2 years

Training background:

9 ± 1 h per week

PB STA:

373 ± 35 s

PB DYN:

133 ± 42 m

EBHD: 148 ± 6 g/L

ND: 149 ± 4 g/L

EBHD: 45 ± 2%

ND: 45 ± 2%

 BHD breath-hold divers, DNF dynamic apnoea no fins, DYN dynamic apnoea with fins, EBHD elite breath-hold divers, ND non-divers, PB personal best, STA static apnoea

*denotes significance (p<0.05) between groups