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. 2021 May 17;12:687495. doi: 10.3389/fphar.2021.687495

TABLE 1.

Clinical and biological features of retinal diseases and confirmed galectin involvement. See main text for additional information.

Diseases of the retina Clinical features Biological features Galectins involved
Physical damage Retinal tear Floaters; Photopsia; Vitreous Haemorrhage; Can result in retinal detachment Landau and Kurz-Levin (2011) Vitreous liquifies with age and eventually detaches from the retina Landau and Kurz-Levi (2011)
Retinal detachment Light flashes, Floaters; Photopsia; Blurred vision; Reduced peripheral vision Landau and Kurz-Levin, (2011) Rhegmatogenous (most common): Caused by hole in the retina. Gal-3 accumulation in subretinal fluid of patients with rhegmatogenous retinal detachment Poulsen et al. (2020)
Tractional (e.g. poorly controlled diabetes): Caused by traction from scar tissue on the retinal surface
Exudative: Not caused by holes or tears, but often by tumours or inflammation Landau and Kurz-Levin (2011); Steelman and Li (2014)
Macular hole Blurring; Distortion Full-thickness defect of retina involving fovea Ittarat et al. (2020)
Proliferative Vitreoretinopathy (PVR) Blinding complication of fibrovascular proliferation Proliferative and inflammatory response of a variety of retinal cells–RPE undergo EMT Alge et al. (2006); Priglinger et al. (2016) Gal-1 and Gal-3 reduce RPE cell adhesion and spreading Alge et al. (2006); Priglinger et al. (2016)
Epiretinal membrane Metamorphopsias and central vision impairment Abnormal growth of tissues on the retinal surface Wang et al. (2016) No direct evidence for galectin, but key role of Gal-3 in fibrosis Slack et al., (2021)
Complex, multifactorial Diabetic Retinopathy (DR) Microaneurysms; Intraretinal haemorrhages; Cotton-wool spots; Venous beading; Vascular loops Degeneration and loss of pericytes; Proliferation of endothelial cells and thickening of the basement membrane; Capillary occlusion and reduced capillary flow; Inflammation; Increased platelet stickiness and aggregation; Increased production of angiogenic factors, especially VEGF Antonetti et al., (2021). Gal-1 upregulated in the vitreous and aqueous humour of PDR patients Abu El-Asrar and Ahmed, (2020);Kanda et al., (2017); Ridano et al., (2017)
Gal-1 is upregulated in retinal tissue of mice with features of DR Kanda et al., (2017)
Gal-1 upregulated in neovascular tufts of OIR mice Liu et al., (2009)
Gal-3 KO mice showed less retinal disease Canning et al. (2007)
Retinal vein occlusion (RVO) Blocked central or branch retinal vein causing widespread retinal haemorrhages and macular oedema (Blair and Czyz (2020)) Neovascular complications Vascular complications similar to DR/DMO, indicating Gal-1 involvement (see main text)
Gal-3 may protect retina as it does in ischemic stroke Wesley et al. (2020)
Retinopathy of prematurity (ROP) Late stages may present with leucocoria (white reflex); Nystagmus with abnormal eye movements; Bilateral retinal detachment; Falciform fold and pthisis bulbi Dogra et al. (2017) Delayed retinal vascular development due to hyperoxia and low serum IGF1 in premature babies; Reflex vasoconstriction; Pathologic angiogenesis; High VEGF in the vitreous Margalit and Srinivas (2003) Galectins not directly involved, but ROP management uses anti-VEGFs, suggesting roles for galectins (see main text)
Hypertensive Retinopathy (HR) Mild or vasoconstrictive (silver or copper wiring) Mild–retinal arterial narrowing of the vessels or sclerosis; moderate–additional intimal thickening and arterial narrowing; focal or diffuse arterial wall opacification Malignant–optic nerve swelling Harjasouliha et al. (2017); Kabedi et al. (2014); Tsukikawa and Stacey (2020)
Moderate or sclerotic phase (hemorrhages, microaneurysms, cotton-wool spots, exudates)
Malignant or exudative phase (moderate retinopathy and optic disk swelling)
Age-related macular degeneration (AMD) Dry (non-neovascular)–slow but progressive decrease in visual acuity, increasing light sensitivity, and reading difficulties Dry–yellow lesions (drusen) below the RPE, atrophy or hyperpigmentation of the RPE Gal-1 upregulated in a model of wet AMD Wu et al. (2019)
Wet (neovascular)–sudden, often quite marked, decrease in visual acuity; can results in permanent reduction of vision as well as a central scotoma Wet–neovascular growth of the choroid; bleeding and exudation from these vessels can damage the outer retina, leading to photoreceptor degeneration Margalit and Srinivas (2003); Landau and Kurz-Levin (2011) Gal-2, -7, -8 upregulated in RPE/choroid samples of some forms AMD; Gal-8, -12 downregulated in neuroretina of pre-AMD patients, and Gal-3 upregulated in most forms of AMD Newman et al. (2012)
Gal-3 upregulated in choroid samples from advanced dry AMD Yuan et al. (2010)
Inherited retinal diseases Retinitis Pigmentosa (RP) Signs include optic nerve pallor, constricted retinal vessels, and bone spicule pigmentation in the periphery Progressive loss of retinal rod photoreceptor cells followed by subsequent degeneration of cones→ increased reduction of retinal function and eventually retinal atrophy Hartong et al. (2006);Landau and Kurz-Levin (2011); Margalit and Srinivas (2003). Gal-3 expression elevated in Müller cells in mouse model of RP Roesch et al. (2012)