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. 2021 Jun 9;14:91. doi: 10.1186/s13045-021-01105-2

Table 3.

Key findings related to malignant cell heterogeneity among tumors obtained using single-cell sequencing

Tumor Technology Platform Key findings References
PDAC scRNA-seq, trajectory analysis 10X Genomics Identified 11 subpopulations of malignant tumor cells, including a subset of malignant ductal cells with unique proliferative features associated with the inactivation of tumor-infiltrating T cells [90]
PDAC scRNA-seq, trajectory analysis 10X Genomics Identified 6 acinar metaplastic cell subpopulations during the progression from preinvasive stages to tumor formation in a mouse model [212]
LUAD, LUSC Bulk and scRNA-seq 10X Genomics Revealed that nongenetic heterogeneity is a major predictor of phenotypic heterogeneity [78]
LUAD scRNA-seq 10X Genomics Identified 19 tumor-specific markers as candidate markers for the detection of extraordinarily rare circulating tumor cells [213]
lung cancer Single-cell sequencing CyTOF Revealed that AXL inhibition suppresses SMAD4/TGFβ signaling and induces JAK1/STAT3 signaling pathways, and AXL functions via CD133-mediated cancer stemness and hybrid EMT features in patients with advanced-stage tumors [214]
GGN-ADC, SADC scRNA-seq, flow cytometry 10X Genomics Revealed the downregulation of signaling pathways associated with angiogenesis and cell proliferation, low expression of collagens of fibroblasts and activated immune cells in GGN-ADC [215]
melanoma scRNA-seq, bulk RNA-seq, ATAC-seq 10X Genomics Confirmed the intermediate state of melanoma cells, which is governed by SOX6, NFATC2, EGR3, ELF1 and ETV4 and regulated by a gene regulatory network [79]
GA scRNA-seq 10X Genomics Identified 5 subtypes of malignant tumor cells, 3 of which corresponded with the histopathological features of Lauren’s subtypes, while 1 was associated with the GA-FG-CCP [216]
EGC scRNA-seq 10X Genomics Revealed that the glandular mucous cells tended to acquire an intestinal-like stem cell phenotype during metaplasia; HSE6 may help identify early-stage tumors [216]
AML scRNA-seq, next-generation sequencing 10X Genomics Identified 3 leukemia subpopulations arrested at different stages of myeloid differentiation: CD34*CD117dim blasts (blocked in G0/G1 phase), CD34*CD117bri blasts and partial maturation myeloid cells [217]
MM scRNA-seq Fluidigm C1 Identified 4 groups of malignant cells; the L1 group from MGUS expressed the lowest levels of genes involved in oxidative phosphorylation, while the L4 group from MM displayed the highest expression [218]
NEPC scRNA-seq 10X Genomics Revealed that focal neuroendocrine differentiation exclusively originates from luminal-like malignant cells and identified differentiation-association signature genes [219]
medulloblastoma scRNA-seq 10X Genomics Identified OLIG2-expressing glial progenitors as transit-amplifying cells at tumor onset, and they are enriched in therapy-resistant and recurrent medulloblastoma [220]
glioma scRNA-seq 10X Genomics Identified Zfp36l1 as a key gene for glioma growth, various transitional intermediate states and corresponding developmental lineages [221]
glioblastoma scRNA-seq 10X Genomics Identified key genes involved in NSC transformation into tumor-promoting cells; NSCs may promote metastasis via extracellular vesicles [222]
glioblastoma scRNA-seq 10X Genomics Identified RAD51AP1 as an independent prognostic factor and a potent mediator of EGFRvlll signaling [223]
ESCC scRNA-seq, bulk sequencing SMART-Seq 2 Identified genes associated with radioresistance, including autophagy-related 9B, DNA damage-inducible transcript 4, myoglobin and plasminogen activator tissue type [224]
HCC scRNA-seq 10X Genomics Identified the involvement of JUNB, a pro-oncogene, in the immune response and progression of HCC [225]
HCC Single-cell mass cytometry, RNA sequencing Fluidigm Revealed that the lncRNA HOXA-AS2 is associated with the regulation of cancer stemness during tumorigenesis and poor prognosis; identified an EPCAM + C-MYC + CK19 subpopulation in HOXA-AS2 high patients [226]
breast cancer scRNA-seq Hydro-Seq Revealed that CAFs increase cancer stemness and alter the epithelial/mesenchymal status during coculture with malignant cells [111]
breast cancer Single-cell whole-exome sequencing Fluidigm C1 Revealed the evolutionary process of cells with SNVs in hit driver genes via CNVs acquired in chromosomal regions; identified the Plekha5 gene as a suppressor of tumor metastasis [227]

PDAC: pancreatic ductal adenocarcinoma; LUAD: lung adenocarcinoma; LUSC: lung squamous cell carcinoma; GGN-ADC: ground glass nodule adenocarcinoma; SADC: solid adenocarcinoma; GA: gastric adenocarcinoma; GA-FG-CCP: fundic gland-type gastric adenocarcinoma; AML: acute myelogenous leukemia; NEPC: neuroendocrine prostate cancer; EGC: early gastric cancer; ESCC: esophageal squamous cell carcinoma; MM: multiple myeloma; HCC: hepatocellular carcinoma; ESCC: esophageal squamous cell carcinoma