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. 2021 Jun 10;16(6):e0252291. doi: 10.1371/journal.pone.0252291

Table 7. Data for swimming flagellated eukaryotes.

Species Geometry Kinematics References
B W Flagella U Notes
Actinomonas mirabilis 240 V = 75. [243]
Alexandrium minutum 21.7 ± 2.2 (64 ± 23 − 320 ± 84) At 12°C. At 24°C. Swimming speeds also available at 8, 16 and 20°C. [244]
Alexandrium ostenfeldii 41.1 ± 4.5 (66 ± 19 − 150 ± 34) At 12°C. At 16°C. Swimming speeds also available at 8 and 20°C [244]
Alexandrium tamarense 26.7 ± 2.6 (108 ± 44 − 255 ± 81) A. tamarense isolate SB50 appeared in doublet configuration and swam at U = 344 ± 52(max472) (compare: single cells U = 238 ± 64(max360)). At 12°C. At 24°C. Swimming speeds also available at 8, 16 and 20°C. [244]
Amphidinium britannicum 51.2 ± 7 (39.5 − 69.9) 36.3 ± 4.6 (30 − 53.9) N = 2 (longitudinal and transverse): LLB, λT = 3.375, hT = 0.74. 72.85(14.1 − 123.3) Algae from Biologische Anstalt Helgoland, ME76, (Herdman) Lebour, measured with Laser Doppler Spectroscopy. [245, 246]
Amphidinium carterae 16.7(10 − 22) 10(8 − 13) L > 7.64. 81.27(14.1 − 149) Algae from Biologische Anstalt Helgoland, ME 30, Hulburt, measured with Laser Doppler Spectroscopy. Average from three ranges registered. Average from two ranges registered. [245, 247250]
Amphidinium klebsi 36.25(20 − 50) 23.25(14 − 30) 73.9 (Synonym of A. operculatum) Average from two ranges registered. [247249]
Apedinella spinifera (6.5 − 10) L = (6.5 − 20). 110 (90 − 175) V = 450, G = (0.5 − 1). [93, 251]
Bodo designis (4 − 7) (2 − 4) N = 2, L1B, L2 ≈ 2B. 39 ± 1(max80) V = 54. [166, 243, 251, 252]
Brachiomonas submarina (15 − 40) (77 − 115) Algae from University of Oslo, Bohlin, measured with Laser Doppler Spectroscopy. [245]
Cachonina niei 21.44 13.36 227.1(50 − 555.6) (Syn.: Heterocapsa niei) Average of the values registered. [253255]
Cafeteria roenbergensis (1.5 − 2.5) (1 − 1.5) N = 2 (longitudinal and transverse):LL = (3 − 5) × B, LT = (1 − 1.5) × B. 103.6 (58 − 131.8)× C. minuta. [251, 256]
Ceratium cornutum (114.5 − 130) (50.7 − 77) (125 − 230.5) Ω = (0.5 − 0.67), ω = 50. [253, 257259]
Ceratium furca (35 − 210) 27.5 N = 2 (longitudinal and transverse). For the longitudinal flagellum: L = (1.97 − 2.2) × W, producing planar sine waves with nw ≈ 1.5, λ = (0.77 − 1.37) × W, h = (0.177 − 0.25) × W. (166 − 222) Measured at 18-20°C. [92, 155, 260262]
Ceratium fusus 240(15 − 600) (15 − 30) N = 2, L = 200, helical or planar beat. (62.5 − 250) 5 measures of speed at 18-20°C. Average from three ranges registered. [91, 92, 155, 260, 263]
Ceratium hirundinella (95 − 700) (194.4 − 277.8) [155, 253, 263]
Ceratium horridum (200 − 250) (40 − 60) (8.3 − 33.3) At 18-20°C, 4 measures of speed. [92, 155]
Ceratium lineatum 82.1 26.8 λ = 18.6, h = 1.3. 36 Ω = 0.63. Cells swim in a helicoidal path with hpath = 6.5 and λpath = 380, with ωormal = 0.07. From illustration. [264]
Ceratium longipes 210 (51 − 57) 166 At 18-20°C, 1 measure of speed. [92, 263]
Ceratium macroceros (40 − 60) 15.4 At 18-20°C, 1 measure of speed. [92, 155]
Ceratium tripos 158(79.56 − 225) (68.1 − 157.1) N = 2 (longitudinal and transverse). For the longitudinal flagellum: L = 224 ± 27, nw = 2.27 ± 0.33, λ = 74.3 ± 9.6, h = 14.2 ± 2.3. Helical or planar beat. 121.7 ± 26.8 (69.4 − 250) ω = 30.2 ± 2, at 20°C and pH8. Average of 4 values registered. Horn length not included. From 6 values registered. [91, 92, 245, 257, 263, 265]
Chilomonas paramecium 32.5 (20 − 40) (10 − 12) N = 2, nw = 1.5. Helical beat. Mastigonemes. 132.35 (59.7 − 162.8) At 25-26°C. From 4 values registered. [91, 247, 263, 266, 267]
Chlamydomonas moewusii (9 − 16) (5 − 12) N = 2, L = (12 − 24). 128 Breaststroke swimming with ω = (10.5 − 12.2). [247, 268, 269]
Chlamydomonas reinhardtii (7 − 13) N = 2, L = (10 − 12). Breaststroke beat. 92.27 (60 − 200) Ω = (0.13 − 0.382), ω = 53 ± 5. Wild-type C. reinhardtii (cc1690). [119, 247, 270273]
Chlamydomonas sp. 13 N = 2, L = 35(≈2.5 × B), nw = 1, λ = 6.3, planar DDW or rowing breaststroke. (61 − 65.4) ω = 8. [19, 91, 245, 247, 274, 275]
Codonosiga botrytis 15 5 N = 1, L = 30. c = 500, ω = 28. [276278]
Crithidia deanei 7.4 ± 0.2 L = 13.1 ± 0.4, nw = 0.77(= L/Λ), λ = 11.7 ± 0.2, h = 2.2 ± 0.05. 45.6 ± 1.5 c = 466 ± 12, ω = 40.5 ± 0.8. Cells were cultured at 28°C, and examined at room temperature, 22°C. [279]
Crithidia fasciculata 11.1 ± 0.3 L = 15.1 ± 0.5, nw = 0.94(= L/Λ), λ = 11.6 ± 0.2, h = 2.2 ± 0.07. 54.3 ± 2.6 c = 680 ± 28, ω = 60 ± 2.3. Cells were cultured at 28°C, and examined at room temperature = 22°C. [279]
Crithidia oncopelti (8 − 8.2) (2.6 − 3) N = 1, L = (17 − 20), λ = 14.4, h = 2.4, planar BDW or DDW. (17 − 20) c = 250, ω = 16.8, Ω = (1 − 2). λ, h and ω are available in function of the pressure of the fluid. [19, 91, 247, 270, 280282]
Crypthecodinium cohnii (101 − 144.6) Data from helical tracking. [264]
Diaphanoeca grandis 40 V = 74. [243]
Dinophysis acuta 65 55 N = 2, L = 65, h = 11. Mastigonemes. Either helical or planar beat of flagella. 500 At 18-20°C, 1 measure of speed. [92, 276]
Dinophysis ovum 45 34 160 [95]
Distigma sp. (43.8 − 105.8) N = 2, L1 ≈ 2B, L2 < B/2, DDW [260, 275]
Dunaliella sp. (8 − 13.5) (5 − 7.6) N = 2. (121 − 226) At 20.5-21.5°C. [91, 96, 245, 247, 263]
Euglena gracilis (45 − 50) (9.2 − 15) N = 1, L ≈ 45. 100.9(59.7 − 162.8) ω = 41.15, Ω ≈ 1.25. Average of three registered values. From video. [91, 247, 263, 266, 283]
Euglena viridis (52 − 64) (10 − 17) N = 1, L = (100 − 128), nw ≈ 1.5, λ = 35 ± 5, h = 6 ± 1, helical DDW. 80 ± 15 (max168) c = (410 − 813), Ω ≈ 1, ω = 12 ± 3. Estimate. [19, 91, 247, 263, 270, 275, 281, 284, 285]
Eutreptiella gymnastica (17 − 30) N = 2, L1 = (20 − 32), L2 = (8 − 13). 240 (200 − 275) G = (2 − 5), V = 650. [93, 251]
Eutreptiella sp. R (40 − 60) (13 − 17) (115 − 155) G = (1.5 − 2.5), V = (4000 − 5888). [93, 155]
Exuviaella baltica (9 − 22) 138.9 (Taxonomic synonym of Prorocentrum balticum) [155, 286, 287]
Giardia lamblia (10.4 − 12.1) (7.3 − 8.9) N = 8 or four pairs (anterior, posteolateral, ventral and caudal), L = (10.6 − 12.5), nw = 2, λ = (2.73 − 5.5), h = (0.2 − 0.31). (12 − 40) ωa = (17-18), ωv = (8-11). Values obtained from curve fitting. Simulated values, for both anterior and posterolateral flagella, η = 1. The cells were attached to the glass slide. [288290]
Gonyaulax polyedra 39.2 ± 3.7 (max 48) 33.3 ± 3.5 (max 45) N = 2, either helical or planar beat. (250 − 278) Ω = 0.65. V = 25700. At 20°C. [243, 247, 263, 291, 292]
Gonyaulax polygramma (30 − 54.1) 46.2 500 [155, 253, 293]
Gymnodinium aureolum 394 [5]
Gymnodinium sanguineum 47.6 ± 4 30.9 ± 3.3 (135.4 − 305.6) Ω = 0.19. V = 16700. At 20°C. [243, 253, 292]
Gymnodinium simplex (7.2 − 14) (6 − 10) (234 ± 34 − 879 ± 39) Strain CCMP 418. Motile behaviour was studied in the presence of the ciliate predator Mesodinium simplex. Approaching swimming. Escaping swimming. [155, 294, 295]
Gyrodinium aureolum (27 − 34) (33 − 245) G = (2 − 3). Algae from University of Oslo, Hulburt, measured with Laser Doppler Spectroscopy. V = 250. [93, 245]
Gyrodinium dorsum 37.5 ± 4.1 31.3 ± 3 N = 2, either helical or planar beat. 324 ± 43.8 (254 − 454) Ω = (1.5 − 2.32). At 20°C. Swimming speed for cells with: short flagella = 240 ± 47(120 − 316) μm s−1, no longitudinal flagella = 147 ± 28.5(93 − 224) μm s−1. [19, 247, 253, 291, 292, 296]
Gyrodinium dorsum 34.5 24.5 N = 1. Planar beat. 148.35 Ω = 8.2 (This Gyrodinium has no helical flagella). [19, 296]
Hemidinium nasutum 26.8 (24.4 − 30) 17.1 (15.2 − 20) 105.6 Ω = 1. [253, 258, 297]
Hemiselmis simplex (4 − 6.5) 3 N = 2. 260 (200 − 450) G = (7 − 10), V = 10. [93, 155, 251]
Heterocapsa pygmea (12 − 15) 10.02 ± 0.74 (89 − 115.7) Algae from Biologische Anstalt Helgoland, ME71, Loeblich et al., measured with Laser Doppler Spectroscopy. [245, 298]
Heterocapsa rotundata (10 − 15) (5 − 10) (102 ± 34 − 564 ± 14) Strain K-483, SCAP motile behaviour was studied in the presence of the ciliate predator Mesodinium simplex. Approaching swimming. Escaping swimming. [155, 294, 299]
Heterocapsa triquetra 17 97 ± 2 Equivalent spherical diameter. [166]
Heteromastix pyriformis (5 − 7) N = 2, L1 = (4 − 5), L2 = (1.5 − 2) × B (Also L1 = 3 × B and L2 = 2 × B). 85 (75 − 100) G = (1.5 − 3), V = 13. (Synonym of Nephroselmis pyriformis). [93, 251, 260, 300]
Hymenomonas carterae (10 − 15) (61 − 113) Coccolithophorid from Biologische Anstalt Helgoland, ME72, (Braarud & Fagerl.) and from the University of Oslo, (Braarud & Fagerl.), measured with Laser Doppler Spectroscopy. [245]
Jakoba libera 19 V = 75. [243]
Katodinium rotundatum (7.5 − 14) (6 − 8) N = 2 (longitudinal and transverse flagellum). 370 (300 − 550) G = (5 − 10). Paulmier 1992, Throndsen 1969 and Campbell 1973. V = (350 − 530). (Taxonomic synonym of Heterocapsa rotundata (Lohmann) G. Hansen) [93, 243, 253, 301]
Leishmania major 12.5 ± 0.3 L = 16.4 ± 0.6, nw = 0.91(= L/Λ), λ = 11.9 ± 0.3, h = 2.9 ± 0.07. 36.4 ± 2 c = 291 ± 4, ω = 24.5 ± 0.8. Friedlin strain V, cultured at 28°C, examined at room temperature = 22°C [279]
Menoidium cultellus 45 7 N = 1, L = 10, nw = 1, λ = 10 ± 2, h = 3 ± 0.5. Helical beat. Mastigonemes. (80 − 193.5) c = 411.7, Ω ≈ 1, ω = 17 ± 3. [267, 284]
Menoidium incurvum (24 − 26) N = 1, LB, nw < 1. Helical DDW. 50 c = 312 (estimate), Ω ≈ 1, ω = 12. [247, 275]
Micromonas pusilla (1 − 3) 90(17 − 100) Algae from the University of Oslo, (Butcher) Parke & Manton, measured with Laser Doppler Spectroscopy. V = 1.5. [93, 245]
Monas stigmata 6 N = 2, L1 = 3, L2 = 15. Planar beat. 269 ω = 47.75. In a 3 mm deep chamber. When between thin slides, one has measured U = 10 and ω = 19. [247, 274]
Monosiga sp. 25 V = 20. [243]
Monostroma angicava (5.9 ± 0.09 − 7.53 ± 0.05) (2.96 ± 0.03 − 3.70 ± 0.03) N = 2, L = (13.21 ± 0.17 − 14.21 ± 0.12). (158.4 − 182.7) In water at 5°C. Male gametes. Female gametes. The planozygotes swam with velocity U = 250.8 at 5°C. [302]
Nephroselmis pyriformis (4.5 − 5) (138 − 189) Algae from the University of Oslo, (N. Carter) Ette, measured with Laser Doppler Spectroscopy. [245]
Oblea rotunda 20 20 420 [95]
Ochromonas danica (6.67 − 10.75) (5.5 − 5.7) λ = 4.5 ± 0.2, h = 0.96 ± 0.12. Leading flagellum with mastigonemes, producing DDW. 77 ± 2 ω = 59 ± 2. At 20°C and η = 1. Data also for η = {2.3, 3.7, 5.6, 7.5, 10}. [28, 303]
Ochromonas malhamensis 3 N = 1, nw = 2.8, λ = 7, h = 1. Mastigonemes. Planar DDW. (55 − 60) ω = 68.44. At 18°C. [19, 91, 285]
Ochromonas minima (3.5 − 6.5) N = 2, L1 = (1 − 2) and L2 = B/3. 75 G = (0.25 − 1.25), V = 25. [93, 251]
Olisthodiscus luteus (15 − 30) 140 (20 − 160) G = (0.5 − 1). Algae from the University of Oslo, N. Carter, measured with Laser Doppler Spectroscopy. V = 600. [93, 245]
Oxyrrhis marina (28.2 − 50.8) N = 2 (longitudinal and transverse). 300 ± 134 (90 − 700) Ω = 9.3. Cells swim in a helicoidal path with hpath = 18 and λpath = 108, with ωnormal = 9.8. Speed increased slightly in the presence of food cells. Average of 7 registered values. [5, 260, 264, 304]
Paragymnodinium shiwhaense 10.9 ± 0.4 (8.4 − 15.2) 8.6 ± 0.3 (5.2 − 11.6) N = 2 (longitudinal and transverse), LL = 10.12, λT = (1.4 − 1.7), hT = (0.81 − 0.92). 571 For cells growing photosynthetically and starved for 2 days. Cells fed with A. carterae were bigger. From illustration. [5, 305]
Paraphysomonas imperforata 42 V = 212. [243]
Paraphysomonas vestita 14.7 nw > 2, h = (1.5 ± 0.3 − 2.6 ± 0.5). Mastigonemes. Complex 3D beat. 70(67.7 − 166) ω = 49 ± 4. At 20-25°C. When a particle made contact with the flagellum, the pattern of flagellar beat changed to a hooked wave and the frequency increased to 74 ± 9 s−1. V = 190. Cells swimming in a circular path. Cells swimming in a helical path. [243, 306, 307]
Pavlova lutheri (5 − 8) (121 − 131) Algae from Biologische Anstalt Helgoland, ME52, (Droop) Green, measured with Laser Doppler Spectroscopy. [245]
Peranema trichophorum 55 (20 − 70) 12 N = 1, L = (40 − 100). Mastigonemes. Tractellar, helical BDW. 20 c = 200, ω = (5 − 6). [19, 91, 247, 263, 270, 275, 281, 308]
Peridinium bipes 42.9 37 291 Ω = 4.99. Cells swim in a helicoidal path with hpath = 17.8 and λpath = 289, with ωnormal = 1.92. [264]
Peridinium cf. quinquecorne (16 − 22) 1500 V = 140000. [243, 245, 253, 309]
Peridinium cinctum 46 (40 − 55) 44 (40 − 200) Ω = 0.83. Algae from Biologische Anstalt Helgoland, ME24, (O. F. Müller) Ehrenb., measured with Laser Doppler Spectroscopy. [245, 253, 258]
Peridinium claudicans (50 − 105) (48 − 75) 215(125 − 333) V = 110000. At 18-20°C, 6 measures of speed. (Taxonomic synonym of Protope-ridinium claudicans) [92, 155, 243, 310]
Peridinium crassipes 102 77 100 V = 204000. At 18-20°C, 1 measure of speed. (Taxonomic synonym of Protope-ridinium crassipes) [92, 243, 311]
Peridinium foliaceum 30.6 ± 3.3 30.6 ± 3.3 185.2 Ω = 2. At 20°C. [292]
Peridinium gregarium (30 − 35) B N = 2, LT ≈ 200. (777.8 − 1805.6) (Taxonomic synonym of Bysmatrum gregarium). Estimate. [253, 312]
Peridinium ovatum (54 − 68) 188(125 − 250) V = 110000. At 18-20°C, 2 measures of speed. (Taxonomic synonym of Protope-ridinium ovatum Pouchet). [92, 243, 313, 314]
Peridinium penardii (25.1 − 32.5) 417 (Taxonomic synonym of Peridiniopsis penardii (Lemmermann) Bourrelly). [315317]
Peridinium pentagonum 106 (75 − 110) 87.5 (75 − 100) 252 (200 − 333) V = 110000. At 18-20°C, 2 measures of speed. (Taxonomic synonym of Protope-ridinium pentagonum (Gran) Balech). [92, 243, 318, 319]
Peridinium subinerme (40 − 60) (40 − 50) (278 − 285) V = 50000. At 18-20°C, 1 measure of speed. (Taxonomic synonym of Protoperidinium subinerme (Paulsen) Loeblich III). [92, 155, 243, 320]
Peridinium trochoideum (20 − 30) (15 − 23) (36 − 70) [253, 316]
Peridinium umbonatum 28(25 − 35) 23(21 − 30) 250 Ω = 1.67. [253, 258, 316]
Phaeocystis pouchetii (4.5 − 8) N = 2, L = 1.5 × B. (21 − 155) Algae from Biologische Anstalt Helgoland, ME64, (Harlot) Lagerh., measured with Laser Doppler Spectroscopy. [245, 251]
Polytoma uvella 18.25(15 − 30) (9 − 20) N = 2, LB, λ = 15, h = 2.9. Planar DDW and rowing breaststroke beating. 103.7(74.8 − 127) c > 312, Ω = (3 − 4), ω = 11.7 (7 − 20). At 20-22°C. Average of the different values registered. [19, 91, 247, 263, 274, 275, 321, 322]
Polytomella agilis (9.8 − 15) (4.9 − 9) N = 4, L = (8 − 9), Planar DDW and rowing breaststroke. (80 − 220) c = (90 − 450), ω = (7 − 33). At 20-22°C. [19, 91, 247, 270, 322, 323]
Poteriodendron sp. L = 35, λ = 4, h ≈ 2. Planar beat. Sessile ω = 40. At 20°C. [281]
Prorocentrum mariae-lebouriae 14.8 ± 1.7 14.8 ± 1.7 (83 − 171.3 ± 27.8) Ω = 3. At 20°C. [245, 257, 292]
Prorocentrum micans (40 − 50) 117.55(47.2 − 611) V = 34000. Average of 6 registered values. [243, 245, 253]
Prorocentrum minimum 15.1 ± 0.3 (max20) 11.8 ± 0.8 N = 2 (longitudinal and transverse): λL = 12.22 ± 0.81, hL = 1.31 ± 0.2, hT = 1.14 ± 0.14. 107.7 ± 54.6 Ω = 1.12 ± 0.23, ωL = 65.9 ± 9.4, ωT = 36.1 ± 15.2. Algae from Biologische Anstalt Helgoland, ME 3, Pavillard, measured with Laser Doppler Spectroscopy. Strain NIES-238 cultured in ESM medium at 20°C. [245, 257]
Prorocentrum redfieldii 33.2 10.28 L = 13.5. 333.3 Bursa (Taxonomic synonym of Prorocentrum triestinum J. Schiller). [324, 325]
Protoperidinium depressum 132 116 450 [95]
Protoperidinium granii (35 − 80) (25 − 56) 86.1 (Ostf.) Balech. [155, 324, 326]
Protoperidinium pacificum 54 50 410 [95]
Prymnesium parvum 7.2 ± 0.3 5.4 ± 0.5 N = 2 and a haptonema. L = 10, Lh = 3.4 ± 0.6. 30 ω = 40. [327]
Prymnesium polylepis 9.1 ± 0.8 6.8 ± 0.4 N = 2 and a haptonema. nw = (1 − 2), L = 28, Lh = 13.5 ± 1.3, λ = 13.4 ± 2.4, h = (1.9 − 2.4). 45 ω = 33.3. [327]
Pseudopedinella pyriformis (5 − 8) N = 1 (and usually a pseudopodium), L = (3 − 5) × B. 105 (90 − 110) G = (0.5 − 1), V = 500. [93, 251]
Pseudoscourfieldia marina (3.2 − 5) (21 − 63) Algae from the University of Oslo, (Throndsen) Manton, measured with Laser Doppler Spectroscopy. [245]
Pteridomonas danica 5.54 Sine waves with λ = 13.0 ± 1.5, h = 2.2 ± 0.5. Mastigonemes. (120.8 − 238.1) ω = 30 ± 4. At 20-25°C. For cells swimming in helicoidal paths. For straight: U = (64.3 − 69.7); and for circular: U = (112 − 134.3). [306, 328]
Pyramimonas amylifera (18 − 31) N = 4 or 8, L = 1.5 × B. (20 − 25) Algae from the University of Oslo, Conrad, measured with Laser Doppler Spectroscopy. [245]
Pyramimonas cf. disomata (6 − 12) (4 − 5) N = 4, L = 8. 350 (290 − 420) G = (4 − 6), V = 100. Using the width given to construct scale bars; average of the four flagella. [93, 251, 329, 330]
Rhabdomonas spiralis (14 − 40) 10 N = 1, nw = 1, λ = 15 ± 3, h = 3.5 ± 0.5. Helical beat. Mastigonemes. 120 ± 20 Ω ≈ 1.4, ω = 25 ± 5. [91, 267, 284]
Rhodomonas salina (12 − 17) 6 N = 2, L ≈ 0.7 × B. (153 ± 16 − 950 ± 90) Strain from the Marine Biological Laboratory, University of Copenhagen. Motile behaviour was studied in the presence of the ciliate predator Mesodinium simplex. Approaching swimming. Escaping swimming. [5, 155, 251, 294]
Scrippsiella trochoidea 25.3 ± 2.4 (max 35) 19.9 ± 2.1 82 (22.2 − 153) V = 3600. Algae from Biologische Anstalt Helgoland, ME64, (Stein) Loeblich, measured with Laser Doppler Spectroscopy. [243, 245, 292, 324]
Spumella sp. 10 25 ± 2 Equivalent Spherical diameter. (Synonym of Monas O. F. Müller 1773 and of Heterochromonas Pascher 1912) [166, 251]
Teleaulax sp. (12 − 15) N = 2, L ≈ 0.6 × B⋆†. (53 − 56) Behaviour in the presence of the predator Oxyrrhis marina. Prior to encounter with predator: U = (61 − 76), post-encounter: U = (133 − 143). For T. acuta [5, 251]
Tetraflagellochloris mauritanica (3 − 5) (2 − 2.5) N = 4, two short flagella (Ls = (11-12)) and two long flagella (Ll = (33-36)). 300 ± 35(260 − 350) ω = 10 ± 1. During forward swimming (the four rear-mounted flagella beat synchronously, uni-directionally, and perfectly phase-locked behind the cell). During backward swimming (the right and left flagella couples beat asynchronously, alternatively and sequentially every 0.4 s), U = 102 ± 13(85 − 120) and ω = 2.5. The cells are also observed to form colonies of up to 16 cells, for which U = 98 ± 11(83 − 115). [331]
Trachelomonas volvocina 25 L = 50. Series of helical waves. Mastigonemes. [267]
Tritrichomonas foetus 14.63 ± 1.3 6.73 ± 1 N = 4. Forces and torques have been characterised but no swimming velocity is given. [332]
Trypanosoma brucei (11.51 − 26) (1.03 − 3.6) L = 9.04, λ = (1.8 − 3.9), h = (1 − 2.5). Planar BDW loops of varying λ and h. (Using the figures of [333] one gets L = 19.4, nw = (1 − 2), λ = 7.5and h = 1.95). (5 ± 2 − 8 ± 2) Propagation of kinks = (85 ± 18 − 136 ± 7) μm s−1. Ω = 19 ± 3 flips s−1 (each flip ≈180° rotation) at 22°C. Procyclic form. Bloodstream form. [333] measured U = 18.6 ± 5.9(9.7 − 38)× in persistent swimming and ω = 19 when swimming in mouse blood. Motility of the strains ILTat 1.4 and AnTat 1.1 was analysed in the blood from different host mammals. The authors also studied the changes in motile behaviour in response to viscosity changes. [333336]
Trypanosoma congolense (11 − 25) (1.8 − 3.7) N = 1, L = (10.47 − 12.76), nw = (1 − 2), λ = (5.26 − 5.84), h = (0.64 − 0.82). 9.7 ± 5.0(1.8 − 26.0)× ω = (6 − 9). Motility of the strain IL 1180 and KETRI 3827 was analysed in the blood from different host mammals. The authors also studied the changes in motile behaviour in response to viscosity changes. [333]
Trypanosoma cruzi 20 2 N = 1, nw = 3, λ = 3.5, h = 0.5. Planar sine BDW. (40 − 304) ω = (14 − 23). In blood. Flexible body. [19, 91, 337, 338]
Trypanosoma evansi 22(19 − 24) 2.9(2.1 − 3.7) N = 1, L = 18.84, nw = (1 − 2), λ = 9.4, h = 1.8. 16.1 ± 5.5(4.7 − 26)× ω = 15. Motility of the strain KETRI 2479 and KETRI 4009 was analysed in the blood from different host mammals. The authors also studied the changes in motile behaviour in response to viscosity changes. [333]
Trypanosoma vivax 23(18 − 29) 3.4(2 − 3.2) N = 1, L = 19.4, nw = (1 − 2), λ = 10.2, h = 2.8. 29.5 ± 19.4 (4.5 − 109)× ω = (13 − 29). Motility of the strains IL 1392 and IL 2136 was analysed in the blood from different host mammals. The authors also studied the changes in motile behaviour in response to viscosity changes. [333]