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Pore geometry is modulated by nucleotide occupancy. Pentamer complex simulated in apo, ATP-bound, and ADP-bound states with substrate DNA (not shown for easier visualization) as Richardson diagrams (top row) and space-filling representations (bottom row). In the Richardson diagrams, the five Arg110s are shown as spheres to highlight their contribution towards DNA gripping in the ATP-bound state. The pore is less constricted in the apo and ADP-bound states than the ATP-bound state.
