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. 2021 Jun 9;11:670917. doi: 10.3389/fonc.2021.670917

Table 1.

LncRNAs that modulate the response of glioblastoma/glioma to chemotherapy.

lncRNA Samples Cell Lines Target/pathway Function Kaplan Meier Analysis Ref
RP11-838N2.4 U87, U251, U87/TMZ, U251/TMZ, miR-10a, TGFB 1, TGFBR1, Smad-2/3/4 RP11-838N2.4 by inhibiting the functions of miR-10a could increase temozolomide cytotoxic effect in GBM. (32)
MALAT1 Mouse U251, U87, U251/TMZU87/TMZ ZEB1, Snail, SLUG MALAT1 by regulating ZEB1 could decrease the sensitivity of resistant GBM. (15)
MALAT1 Mouse LN-229, LN-428,LN-319, LN-18, p-MKK3/6, p-p38,P-ERK, WNT/Ca2+ WIF1 could increase the migratory possibility of GBM via WNT5A that activates the WNT/Ca2+ pathway and MALAT1. (16)
MALAT1 Human U87, U251, U87/TMZ, U251/TMZ miR-203, TS MALAT1 could induce chemoresistance to TMZ by suppressing miR-203 and promoting thymidylate synthase expression. (17)
MALAT1 Human U251, U251/TMZ miR-101, MRP1, MGMT, p-gp Knockdown of MALAT1 by promoting miR-101 could inhibit resistance to TMZ. (18)
MALAT1 Human,Mouse U87, T98G, LN-18,U87/TMZ, T98G/TMZ, LN-18/TMZ AERG, CCL2, CXCL4 MALAT1 silencing could sensitize glioblastoma to TMZ. (33)
MALAT1 U87, A172, U251, U87/TMZ,A172/TMZ, U87/TMZ p53, NF-kB p50 and p52 are primary regulators of this ncRNA. (19)
TUSC7 Human U87, U87TMZ miR-10a, MDR1 TUSC7 by targeting miR-10a could inhibit TMZ resistance in GBM. (29)
NEAT1 Human U87, U251, U87/TMZ, U251/TMZ let-7g-5p, MAP3K1, E-cadherin, N-cadherin NEAT1 by regulating the let-7g-5p/MAP3K1 axis could promote malignant phenotypes and TMZ resistance in GBM. (20)
NEAT1 Human U87, U87/R, U251, U251/R MGMT NEAT1 by regulating MGMT could be involved in TMZ resistance in GBM multiforme. (21)
H19 Human A172, LN229, U87MG, LN18, T98G NKD1 H19 could contribute to NKD1 repression via the recruitment of EZH2 on its promoter. (22)
H19 Human U87, U251
U87/TMZ
U251/TMZ
PARP, MDR,
MRP, ABCG2
Knockdown of H19 could enhance the sensitivity of human glioma cells to TMZ. (34)
H19 U251, LN229
U251/TMZ LN229/TMZ
Caspase-3, NF-kB H19 By activating NF-kB signaling could confer TMZ resistance in glioma. (23)
H19 U-251, M059J,
U251/TMZ,
M059J/TMZ
Wnt/β-catenin,
Vimentin,ZEB1, c-myc,E-cadherin,
Survivin
H19 silencing by suppressing EMT via the Wnt/β-catenin pathway could reduce the resistance of human glioma cells to TMZ. (11)
UCA1 Human U251,U87MG CXCL4, miR-182, PFKFB2 UCA1/miR-182/PFKFB2 axis could modify GBM-associated stromal cells-mediated glycolysis and invasion of glioma cells. (35)
uc003iax.2, ENST00000443252 Human U87, U251, U87/TMZ, U251/TMZ IL-18, DPP4, ABCB1, TP53, Collagen I, Fibronectin, Laminin Dysregulated lncRNAs could be involved as novel targets so as to overcome acquired TMZ resistance in GBM chemotherapy. (31)
AC023115.3 Human U87MG,U251MG PARP,Caspase-3 AC023115.3 could suppress the chemoresistance of GBM by decreasing autophagy. (36)
AC003092.1 Human U87, U251, U87/TMZ, U251/251 TFPI-2,miR-195 AC003092.1 could help TMZ chemosensitivity via the miR-195/TFPI-2 axis modulation in GBM. (37)
TP73-AS1 Human G26, G7, G26/TMZG7/TMZ ALDH1A1 TP73-AS1 is involved in aggressiveness and could promote TMZ resistance in GBM cancer stem cells. (38)
ADAMTS9-AS2 Human T98G, U118, T98G/TMZU118/TMZ FUS/MDM2, Tubulin ADAMTS9-AS2 by upregulating the FUS/MDM2 ubiquitination axis could help TMZ resistance in GBM. (39)
SNHG15 Human HMC3, HMC3/TMZ miR-627, EGFR, CDK6, Sox2, β-catenin Modulating SNHG15/CDK6/miR-627 axis by palbocicli could reduce M2-polarization of glioma-associated microglia in GBM multiforme and finally could overcome TMZ resistance. (40)
SNHG12 Mouse N3S, N3T3rd, U251, U251T3rd PARP, Caspase-3, RB, CDK4, CDK6, Cyclin-D1, P-MEK, DNMT1, DNMT3a, DNMT3b, MAPK1, E2F7, P-ERK1/2 Knockdown of SNHG12 by increasing MAPK1 and E2F7 expression and activating the MAPK-ERK could restore TMZ sensitivity in GBM. SNHG12 expression is associated with poor prognosis in GBM. (41)
NONHSAT163779 Human U87, U87/TMZ hsa_circ_0043949, MDR1, MRP1, BCRP, MGMT NONHSAT163779 and hsa_circ_0043949 could be involved as prognostic biomarkers for the treatment of GBM. (42)
SBF2-AS1 Human U87, LN229, A172, T98, U251 XRCC4, y-H2AX, Pro-caspase-3, Cleaved-caspase 3 Knockdown of SBF2-AS1 could increase sensitivity to TMZ in GBM. SBF2-AS1 expression is associated with poor prognosis in GBM. (43)
OKN-007 Rat U138, LN18, T98, U251 TGFβ1 OKN-007 could enhance TMZ sensitivity and suppresses TMZ-resistant GBM. (44)
SOX2OT Human U87, U251, U87/TMZ, U251/TMZ MDR1, BCRP1, MRP1, SOX2, ALKBH5, TCF1, Caspase-3/7/8/9, Wnt5a/β-catenin, Cyclin-D1, C-Myc, LEF1 SOX2OT by elevating SOX2 expression via ALKBH5-mediated epigenetic regulation could promote TMZ resistance. Elevated SOX2OT expression is associated with poor prognosis in GBM. (45)
00021 BALB/c U87, U251, A172, and SHG44 P21, Notch1, Hes1, Hes5 Long intergenic noncoding RNA 00021 by epigenetically silencing p21 via the Notch pathway could promote GBM TMZ resistance. LINC00021 expression is associated with the poor prognosis of GBM patients. (46)
MIR22HG Mouse U87MG, LN229, LN1 Wnt/β-catenin, P21, P27, c-Muc, p-GSK3B, Cyclin-D1, LEF1 MIR22HG via suppressing the Wnt/β-catenin pathway could inhibit GBM progression. MIR22HG expression is associated with poor prognosis in GBM (12)
HOTAIR Mouse A172, LN229, A172/TMZ, LN229/TMZ miR-519a-3p, RRM1, Vimentin, E-cadherin, CD63, N-cadherin, MAP3K1 Knockdown of HOTAIR by miR-519a-3p/RRM1 axis could regulate TMZ resistance. (24)
HOTAIR Human U87, A172, U87/TMZ, A172/TMZ miR-125, Cyt C, Caspase-3, HK2 HOTAIR by targeting miR-125 could promote chemoresistance in human GBM. (25)
MIR155HG Mouse, databases A172, U251,
A172/TMZ,
U251/TMZ
Wnt/β-catenin, c-Myc, PTBP1,
Cyclin-D1
Knockdown of MIR155HG by inhibiting the Wnt/β-catenin pathway via downregulation PTBP1 could increase glioma sensitivity to TMZ. MIR155HG Upregulation was associated with poor prognosis (13)
KCNQ1OT1 C Mouse U251, U87,
U251/TMZ,
U87/TMZ
miR-761, c-MYC,
Pim-1L, p-MDR1,
MDR1, Survivin
KCNQ1OT1 C by retrieving PIM1 From
miR-761 could confer gliomas resistance to TMZ.
(47)
NCK1-AS1 Human U251, A172,
U251/TMZ,
A172/TMZ
TRIM1,
miR-137, TRIM24
NCK1-AS1 by modulating the miR-137/TRIM24 axis could increase the resistance of glioma cells to TMZ. (48)
NCK1-AS1 Human A172, LN229
A172/TMZ LN229/TMZ
miR-22-3p, IGF1R NCK1-AS1 via miR-22-3p/IGF1R axis could enhance chemoresistance in glioma. (49)
EPIC1 SNB19, T98G, U97MG,
SNB19/TMZ,
T98G/TMZ,
U97MG/TMZ
Cdc20 Overexpression of EPIC1 via targeting Cdc20 could be useful in glioma treatment. (50)
HOXD-AS1 TCGA dataset U87, U251,
U373, SNB19, U87/DDP, U251/DDP
miR-204, Caspase-3/9 Knockdown of HOXD-AS1 by buffering miR-204 could enhance cisplatin sensitivity. high HOXD-AS1 expression had a poor
prognosis
(51)
LINC01198 Human U251, SNB-19,
LN229, U87,
U87/TMZ,
U251/TMZ,
LN229/TMZ,
SNB-19/TMZ
PTEN, AKT,
NEDD4-1
Overexpression of LINC01198 by enhancing the NEDD4-1-dependent repression of PTEN could promote glioma cell proliferation and resistance to TMZ. LINC01198 high elevation was associated with a poor prognosis of glioma. (52)
LINC00174 Human U251, U87,
U251/TMZ,
U87/TMZ
SOX9,
P13K/Akt
Knockdown of LINC00174 by regulating miR-138-5p/SOX9 axis could decrease chemoresistance to TMZ in glioma. (53)
GAS5 U138, LN18
U87MG, U251MG,
U138/Cis, LN18/Cis, U87/Cis, U251/Cis
mTOR, LC3I,
LC3II, p-62
GAS5 by suppressing excessive autophagy in an mTOR‐dependent manner could facilitate glioma cell sensitivity to cisplatin. (54)
CASC2 Human U257, U87, U257/TMZ, U87/TMZ mTOR, Beclin1,
miR-193a-5p, LC3II/LCI
Upregulation of CASC2 through autophagy inhibition by buffering miR-193a-5p and regulating mTOR expression could sensitize glioma to TMZ cytotoxicity. (55)
CASC2 Human U251, U373, SNB19, U118, LN229
SNB19/TMZ, U251/TMZ
PTEN, AGO2,
Akt, miR-181a
CASC2 by inhibiting miR-181a could increase sensitivity to TMZ in glioma. CASC2 upregulation was associated with poor prognosis (56)
CCAT2 Human U251, U87, A172, SHG44 miR-424 CCAT2 by disturbing the normal function of miR-424 could enhance resistance in glioma. CCAT2 upregulation was associated with a poor prognosis. (57)
DANCR Mouse U87MG, LN18, U251MG, U138MG, U87MG/Cis,
U251MG/Cis, U138MG/Cis
AXL, NF-kB,
IkBa, PI3K/AKT
DANCR via activating AXL/PI3K/Akt/NF-κB signaling pathway could mediate cisplatin resistance in glioma cells. (58)
MEG3 U87, U87/Cis p-62, LC3 I/II,
PARP
MEG3 by suppression of autophagy could enhance cisplatin resistance in glioma. (30)
MSC-AS1 Human LN229, HG-44
LN229/TMZ, SHG-44/TMZ
miR-373-3p, CPEB4, Bax, MCL-1, MRP-1, P-PIK3,
Cyclin-D1,
Caspase-3, PI3K/AKT
Knockdown of MSC−AS1 by regulating miR-373-3p/CPEB4 axis via PI3K/Akt pathway could inhibit cell growth and TMZ resistance in glioma. MSC−AS1 upregulation was associated with a poor prognosis. (50)
NR5A2 Mouse U138, U251, A172, U87, U138/TMZ, U251/TMZ NR5A2, PARP,
NOTCH1, p21,
Cyclin-D1,
caspase-3, MMP2,
E-cadherin
NR5A2 via regulating notch signal pathway could promote cell growth and resistance to TMZ in glioma. NR5A2 overexpression was associated with the poor prognosis of glioma patients (59)
ZFAS1 Human U87, U251, NHA, A172, LN299, LN299/Cis, U251/Cis, miR‐432‐5p Knockdown of ZFAS1 by upregulating miR‐432‐5p could enhance cisplatin cytotoxicity in glioma. Expression levels of
ZFAS1 in clinical tissues is associated with poor prognosis
(60)
XIST Human LN229, U251, LN229/TZM, U251/TZM Ago2, miR-29c XIST via interacting with miR-29c and through DNA mismatch repair pathway could modulate the chemoresistance of glioma cell to TMZ. Higher expression of XIST was associated with a lower OS rate. (61)