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Design and mechanism of the FLIM-MDTP in measuring tensile forces at cell–cell junctions. The FLIM-MDTP can anchor onto cell membranes through a lipid moiety. Intercellular ligand-receptor binding induces the unfolding of a DNA hairpin and results in the separation of a fluorophore-quencher pair, which further leads to an increase in the number of probes in the unquenched state, exhibiting a higher fluorescence lifetime.
