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. 2021 Jul 7;12:705575. doi: 10.3389/fphys.2021.705575

TABLE 1.

Major studies on identifying Sigmar1’s subcellular organelle localization (in chronological order).

Subcellular organelles Experimental methods Cells or Tissues Antibody and antigen Antibody validation References
Mitochondria Microsomal Synaptosomal fractions Western blotting on subcellular fractions Rat brain Anti-Sigmar1 Rabbit Polyclonal antibody raised against synthetic amino acid residues 143–163 of Sigmar1 (produced by authors) Not listed Yamamoto et al., 1999
Plasma membrane Mitochondrial membranes Vesicles or elongated cisternae of ER Immunoelectron microscopy Adult rat brain hypothalamus and hippocampus sections Anti-Sigmar1 Rabbit Polyclonal IgG raised against synthetic peptide of Sigmar1 amino acid sequence 143–162 (produced by authors) Antibody specificity confirmed through absence of staining in brain tissue sections when co-incubated with Sigmar1 specific antigen Alonso et al., 2000
Perinuclear areas Plasmalemmal regions of cell-cell contact Growth cones of neurites Fluorescence microscopy of co-immunocytohistochemistry and co-immunoprecipitation of endogenous Sigmar1 with Ankyrin B and IP3R3 receptors Mouse neuroblastoma x rat glioma hybrid NG-108-15 cells Anti-Sigmar1 Rabbit antibody raised against N-terminal residues of mouse Sigmar1 (produced by authors) Not listed Hayashi and Su, 2001
Mitochondrial membranes i. Radioisotope labeled Sigmar1 agonist [3H](+)-pentazocine binding assay ii. Immunofluorescence i. Isolated rat liver mitochondria ii. Rat liver sections Anti-Sigmar1 Rabbit Polyclonal IgG (Alonso et al., 2000) Validated by original contributors at (Alonso et al., 2000) Klouz et al., 2003
Plasma membranes i. Exogenous expression of N-terminal and C-terminal GFP tagged Sigmar1 constructs and fluorescence microscopy ii. Co-immunoprecipitation with anti-Kv1.4 antibody i. Xenopus Oocytes ii. Triton X-100 solubilized membrane lysates of rat pituitary gland Anti-Sigmar1 Rabbit Polyclonal IgG (Yamamoto et al., 1999) Not listed Aydar et al., 2002
i. ER associated detergent insoluble lipid droplets/microdomains ii. Sigmar1 immunostained reticular structure was noted negative for endosomes (EEA-1), mitochondria (Mitotracker, bcl-2), lysosomes (Lyso Tracker, LAMP-1), synaptic vesicles (Synapsin II), plasma membrane (Fas and CTx-B), golgi (GM130), ER-associated proteins Fluorescence microscopy; cells were specially treated with 0.02% SDS for 10 min NG-108 cells (Hayashi and Su, 2003a, b), Rat hippocampal differentiated oligodendrocytes culture (Hayashi and Su, 2004) i. Exogenously expressed C-terminal Enhanced Yellow Fluorescent Protein (EYFP)-tagged mouse Sigmar1 construct ii. Polyclonal rabbit anti-guinea pig Sigmar1 antibody raised against guinea pig Sigmar1 amino acid sequence 144–165 (Hayashi and Su, 2003b). i. Exogenous overexpression ii. Anti-Sigmar1 staining pattern have been visually compared to expression pattern of exogenously expressed Sigmar1-EYFP plasmid construct (Hayashi and Su, 2003a, b) Hayashi and Su, 2003a, b, 2004
Mitochondria Mitochondria-associated ER membranes (MAM) Nuclear and microsomal fractions i. Immunofluorescence staining of endogenous Sigmar1 in cells transfected with Mitochondria and ER fluorescent constructs, ii. Western blot analysis on ultracentrifugation based subcellular fractionations Chinese hamster ovary (CHO) cells Anti-Sigmar1 Rabbit antibodies raised against amino acid residues 52–69 and 143–165 of Rat Sigmar1 (produced by authors) Western blot on CHO cell lysates and on different rat organs Hayashi and Su, 2007
Focal adhesion contacts (FACs) Co-localization of Sigmar1 with FAC protein Talin in immunofluorescence staining CHO-K1 cells Anti-Sigmar1 Rabbit antibody raised against Maltose Binding Protein (MBP)-Guinea Pig full length Sigmar1 fusion protein (Ramachandran et al., 2007) (produced by authors) Western blot analysis using guinea pig Sigmar1 overexpressing COS-7 cells as positive control (Ramachandran et al., 2007) Mavlyutov and Ruoho, 2007
i. Motor neurons of mouse brain Medulla and spinal cord ii. Cholinergic post-synaptic terminals co-localized to Kv2.1 potassium channels iii. Putative subsurface ER cisternae close to plasma membrane i. Immunohistochemistry ii. Co-immunofluorescence with cholinergic neuron markers iii. Immuno-electron microscopy Mouse brain and spinal cord sections Anti-Sigmar1 Rabbit antibody raised against Maltose Binding Protein (MBP)-Guinea Pig full length Sigmar1 fusion protein (Ramachandran et al., 2007) Sigmar1 knockout mice brain sections as negative control in immunohistochemistry Mavlyutov et al., 2010, 2012
Mitochondria ER Fluorescence microscopy Mouse neuroblastoma Neuro-2a cells Exogenously expressed eGFP and mCherry tagged Sigmar1’s correlative, visual colocalization observation with erRFP and mtGFP labeled ER and mitochondria, respectively Exogenous overexpression Shioda et al., 2012
Mitochondria and MAM fractions Western blot analysis on subcellular fractions Chinese hamster ovary (CHO) cells Anti-Sigmar1 Rabbit antibodies raised against Rat Sigmar1 (Hayashi and Su, 2007) Western blot on CHO cell lysates and on different rat organs (Hayashi and Su, 2007) Mori et al., 2013
ER and Nuclear Envelope Confocal immunofluorescence staining NG108 and Neuro-2a cells Anti-Sigmar1 (Santa Cruz Biotechnology) Commercial vendor Tsai et al., 2015
Predominantly in nuclear envelope, sparsely at ER cisternae located subsurface of plasma membranes Immuno-electron microscopy Mouse retinal photoreceptor cells Anti-Sigmar1 Rabbit antibody raised against Maltose Binding Protein (MBP)-Guinea Pig full length Sigmar1 fusion protein (Ramachandran et al., 2007) Reported at (Ramachandran et al., 2007) Mavlyutov et al., 2015b
Plasma membrane ER Nuclear envelope Immuno-electron microscopy Mouse and rat dorsal root ganglion (DRG) cells Anti-Sigmar1 Rabbit antibody raised against Maltose Binding Protein (MBP)-Guinea Pig full length Sigmar1 fusion protein (Ramachandran et al., 2007) Validated through absence of immunofluorescence staining using Sigmar1 knockout mouse DRG tissue Mavlyutov et al., 2016
Nucleus Nucleoplasmic reticulum Plasma membrane subsurface ER cisternae Immuno-electron microscopy NSC34 cells (produced by fusion of motor neuron enriched, embryonic mouse spinal cord cells with mouse neuroblastoma), Sigmar1-null NSC34 cells Endogenous Sigmar1 stained with antibody as reported in Ramachandran et al. (2007), Exogenously expressed full length Sigmar1-GFP-APEX2 (ascorbate peroxidase 2) fusion construct were enhanced using 3, 3′-diaminobenzidine (DAB) incubation Reported at (Ramachandran et al., 2007) Mavlyutov et al., 2017
i. ER, Mitochondria associated ER membranes (MAM) ii. Cholesterol-containing giant unilamellar vesicles (GUVs) Immuno-fluorescence microscopy HEK293T cells i. Exogenously expressed GFP-tagged human Sigmar1 colocalization assessed with mCherry tagged Sec16β protein and immunofluorescence staining of mitochondrial outer membrane protein Tom20 ii. GFP-tagged Sigmar1 fusion constructs i. and ii. Exogenous overexpression of GFP-tagged fusion constructs Zhemkov et al., 2021