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. 2021 Jul 26;11:15162. doi: 10.1038/s41598-021-94602-w

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© The Author(s) 2021

Open AccessThis article is licensed under a Creative Commons Attribution 4.0 International License, which permits use, sharing, adaptation, distribution and reproduction in any medium or format, as long as you give appropriate credit to the original author(s) and the source, provide a link to the Creative Commons licence, and indicate if changes were made. The images or other third party material in this article are included in the article's Creative Commons licence, unless indicated otherwise in a credit line to the material. If material is not included in the article's Creative Commons licence and your intended use is not permitted by statutory regulation or exceeds the permitted use, you will need to obtain permission directly from the copyright holder. To view a copy of this licence, visit http://creativecommons.org/licenses/by/4.0/.

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The extracellular domain of human ACE2 and RBD of the SARS-CoV-2 S protein are shown with gray and purple respectively. The RBD attached O-glycan is shown with atom name sticks. N-glycans in the proximity of RBD-ACE2 interface are shown with orange spheres. Persistent polar interactions between H1 and H2 of ACE2 and the attached O-glycan are shown in A-C.