Skip to main content
. 2021 Jan 30;70(5):1015–1032. doi: 10.1093/sysbio/syab004

Table 2.

Computational complexity of the proposals, likelihood operations and condition under which a proposal is strictly less expensive than a likelihood evaluation (i.e., Cost condition).

Operation Complexity Cost condition
Full lik. Inline graphic
Partial lik. Inline graphic
Partial pars. Inline graphic
stNNI Inline graphic Inline graphic
eSPR Inline graphic Inline graphic
eTBR Inline graphic Inline graphic
G-stNNI Inline graphic Inline graphic
G-NSPR Inline graphic Inline graphic
A-stNNI Inline graphic Inline graphic
A-2SPR Inline graphic Inline graphic
A-PBJ Inline graphic Inline graphic
  Inline graphic  

Notes: Alignments have Inline graphic sites for Inline graphic sequences defined over an alphabet of Inline graphic characters (e.g., Inline graphic for nucleotide sequences). The Gamma model for rate heterogeneity has a number Inline graphic of rate categories (Yang 1994), while a partial likelihood requires Inline graphic operations instead of Inline graphic. Finally, Inline graphic represents the number of splits altered by a move. For the A-PBJ proposal, Inline graphic, Inline graphic and Inline graphic identify the number of edges in the two unstable paths and stable path, respectively.