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. 2021 Jul 29;12(8):1169. doi: 10.3390/genes12081169
GC1 GC-content at the first codon positions
GC2 GC-content at the second codon positions
GC12 The average of GC1 and GC2
GC3 GC-content at the third codon positions
GC3s Frequency of either a G or C at the third codon position of synonymous codons