Comparison of type I ABC transporters. (A) ABC methionine transporters in the inward-facing conformation. (B) ABC molybdate transporters in the inward-facing conformation and (C) ABC methionine transporters in the outward-facing conformation. For each model the TMDs, NBDs, and SBPs, are colored in blue, grey, and pink, respectively. Next to each panel, an overlay is included of the two structures, with the ABC transporter with a regulatory domain colored in grey and the ABC transporter with a regulatory domain colored in red. NBD:NBD inter-subunit distances were assessed using the Cα of NBD glycine residues in the P loop and signature motifs. The average of the two distances are indicated by double arrows. (D) Schematic summary for NBD:NBD distances of homologous methionine ABC transporters with and with out the autoinhibitory domain in the inward and outward-facing conformations, respectively.