There are two mechanisms by which macrophages release SQSTM1 into the extracellular environment (active secretion or passive release). Extracellular LPS induces SQSTM1 lysosomal secretion by activating the TLR4-MYD88-STING1 pathway, while intracellular LPS triggers the pyroptotic release of SQSTM1 by activating the CASP11-GSDMD-GSDMD-N pathway [78]. After its release, SQSTM1 binds to the receptor INSR to activate the NFκB pathway, causing the polarization of pro-inflammatory macrophages (aka ‘M1’), and finally mediating septic death in mice through excessive inflammation and coagulation [78]. Abbreviations: CASP11, caspase 11; GSDMD, gasdermin D; INSR, insulin receptor; LPS, lipopolysaccharides; MYD88, MYD88 innate immune signal transduction adaptor; NFKB, nuclear factor-κB; PLCG1, phospholipase C gamma 1; ROS, reactive oxygen species; STING1, stimulator of interferon response CGAMP interactor 1; SQSTM1, sequestosome 1; TLR4, toll like receptor 4.