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. 2021 Sep 16;11:17311. doi: 10.1038/s41598-021-96754-1

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© The Author(s) 2021

Open Access This article is licensed under a Creative Commons Attribution 4.0 International License, which permits use, sharing, adaptation, distribution and reproduction in any medium or format, as long as you give appropriate credit to the original author(s) and the source, provide a link to the Creative Commons licence, and indicate if changes were made. The images or other third party material in this article are included in the article's Creative Commons licence, unless indicated otherwise in a credit line to the material. If material is not included in the article's Creative Commons licence and your intended use is not permitted by statutory regulation or exceeds the permitted use, you will need to obtain permission directly from the copyright holder. To view a copy of this licence, visit http://creativecommons.org/licenses/by/4.0/.

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Large-sized Proboscipeda tracks attributed to P. antiquus adults. (a) to (d) PAT/MTS/001: Right manus showing clearly 5 toe impressions and the frontal and lateral displacement rims (morphological interpretation based on the orthogonal (b) and oblique (d) depth and contour (c) maps). (e) and (f) PAT/MTS/010e: Deeper manus with pockmarks; toe pad impressions indicated (I–III). (g) PAT/MTS/004a,b: large manus-pes couple where the hind foot deformed the fore foot during overstepping, and revealing a typical elephantine gait; the toe impressions in both tracks indicate the direction of movement. Scale bar = 10 cm.