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. 2021 Sep 22;11:18863. doi: 10.1038/s41598-021-98409-7

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© The Author(s) 2021

Open Access This article is licensed under a Creative Commons Attribution 4.0 International License, which permits use, sharing, adaptation, distribution and reproduction in any medium or format, as long as you give appropriate credit to the original author(s) and the source, provide a link to the Creative Commons licence, and indicate if changes were made. The images or other third party material in this article are included in the article's Creative Commons licence, unless indicated otherwise in a credit line to the material. If material is not included in the article's Creative Commons licence and your intended use is not permitted by statutory regulation or exceeds the permitted use, you will need to obtain permission directly from the copyright holder. To view a copy of this licence, visit http://creativecommons.org/licenses/by/4.0/.

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Mitochondria in stressed cones expel electron-dense, membranous contents through the plasma membrane. (A,B) Electron dense membranous aggregates in mitochondria are associated with electron-lucent domains of the mitochondrion inter-membrane space (white arrows). (C–H) Mitochondria engage with the plasma membrane, undergo a series of membrane fusions and eject the membranous aggregate into the inter photoreceptor space. (White arrows: swollen or ruptured mitochondrial outer membrane, black arrows: ruptured plasma membrane.